UPSC MainsBOTANY-PAPER-II201115 Marks
Q9.

Comment on EPSPS and Tra genes.

How to Approach

This question requires a detailed understanding of EPSPS and Tra genes, particularly in the context of herbicide resistance in genetically modified crops. The answer should define both terms, explain their roles in plant physiology and genetic engineering, and discuss the implications of their use. Structure the answer by first defining EPSPS, then explaining the Tra genes and their function, followed by a discussion of the benefits and concerns associated with these technologies. Focus on the scientific principles and their application in agriculture.

Model Answer

0 min read

Introduction

The increasing global population demands higher agricultural productivity, leading to the widespread adoption of genetically modified (GM) crops. A key component of many GM crops is the introduction of genes conferring herbicide tolerance, enabling farmers to control weeds effectively. Two crucial elements in this process are 5-enolpyruvylshikimate-3-phosphate synthase (EPSPS) and Tra genes. EPSPS is an enzyme essential for plant growth, while Tra genes encode for proteins that detoxify herbicides. Understanding these genes is vital for comprehending the science behind herbicide-tolerant crops and their impact on agriculture and the environment.

EPSPS: The Target of Glyphosate

5-enolpyruvylshikimate-3-phosphate synthase (EPSPS) is an enzyme crucial in the shikimate pathway, which is responsible for the biosynthesis of aromatic amino acids (phenylalanine, tyrosine, and tryptophan) in plants, bacteria, and fungi. Animals lack this pathway, making EPSPS an attractive target for herbicides. Glyphosate, a broad-spectrum herbicide, inhibits EPSPS, thereby disrupting aromatic amino acid synthesis and ultimately killing the plant.

Naturally occurring EPSPS is sensitive to glyphosate. However, certain bacteria possess EPSPS variants that are less susceptible to glyphosate inhibition. These glyphosate-resistant EPSPS genes have been identified and introduced into crop plants, conferring tolerance to glyphosate. The most commonly used glyphosate-resistant EPSPS gene is derived from the bacterium Agrobacterium tumefaciens CP4.

Tra Genes: Detoxifying the Herbicide

Tra genes (tolerance-related genes) are another mechanism employed to confer herbicide tolerance. Unlike EPSPS modification, Tra genes encode enzymes that actively detoxify the herbicide, breaking it down into non-toxic compounds. These genes are often sourced from bacteria that have evolved mechanisms to survive in herbicide-rich environments.

The most prominent example is the CP4 EPSPS gene coupled with the Tra gene encoding glyphosate oxidoreductase (GOX) from Pseudomonas fluorescens. GOX catalyzes the oxidation of glyphosate, rendering it harmless to the plant. This dual approach – a modified target site (EPSPS) and a detoxification mechanism (Tra) – provides a robust level of glyphosate tolerance.

Comparison of EPSPS Modification and Tra Gene Approach

Feature EPSPS Modification Tra Gene Approach
Mechanism Altered target site; reduced herbicide binding affinity Herbicide detoxification; breakdown into non-toxic compounds
Gene Source Bacteria (e.g., Agrobacterium tumefaciens) Bacteria (e.g., Pseudomonas fluorescens)
Effectiveness Provides tolerance by reducing herbicide impact Provides tolerance by eliminating herbicide activity
Potential for Resistance Development Higher potential for weed resistance due to selection pressure on the target site Lower potential for resistance development as herbicide is degraded

Applications and Concerns

Crops engineered with EPSPS and Tra genes, such as glyphosate-tolerant soybeans, corn, cotton, and canola, have become widely adopted globally. These crops simplify weed management, reduce tillage (leading to soil conservation), and potentially lower herbicide use in some cases. However, concerns exist regarding the development of glyphosate-resistant weeds, the potential for gene flow to wild relatives, and the environmental impact of increased herbicide application in certain regions. The emergence of glyphosate-resistant weeds is a significant challenge, requiring farmers to use more potent herbicides or revert to traditional weed control methods.

Furthermore, the widespread use of glyphosate has raised concerns about its potential human health effects, although regulatory agencies like the EPA and EFSA have consistently concluded that glyphosate is unlikely to pose a carcinogenic risk to humans when used according to label instructions. Ongoing research continues to evaluate the long-term impacts of glyphosate and glyphosate-tolerant crops.

Conclusion

EPSPS and Tra genes represent significant advancements in agricultural biotechnology, enabling the development of herbicide-tolerant crops that have contributed to increased food production. However, the benefits must be carefully weighed against the potential risks, including the evolution of herbicide-resistant weeds and environmental concerns. Sustainable agricultural practices, integrated weed management strategies, and continued research are crucial for maximizing the benefits of these technologies while minimizing their negative impacts. A balanced approach is essential to ensure food security and environmental sustainability.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Shikimate Pathway
A metabolic pathway used by plants, bacteria, and fungi for the biosynthesis of aromatic amino acids, vitamins, and other essential compounds. It is absent in animals, making enzymes in this pathway attractive herbicide targets.
Gene Flow
The transfer of genetic material from one organism to another, particularly from genetically modified crops to wild relatives, potentially leading to the spread of herbicide tolerance traits in unintended populations.

Key Statistics

In 2022, approximately 95% of soybeans and 70% of corn grown in the United States were genetically engineered to be herbicide-tolerant, primarily to glyphosate.

Source: USDA Economic Research Service (ERS), 2023

The global glyphosate market was valued at USD 11.75 billion in 2023 and is projected to reach USD 14.48 billion by 2032.

Source: Fortune Business Insights, 2024

Examples

Roundup Ready Crops

Monsanto (now Bayer) developed "Roundup Ready" crops, which are genetically engineered to express the CP4 EPSPS gene, conferring tolerance to Roundup, a glyphosate-based herbicide.

Frequently Asked Questions

What is the difference between first-generation and second-generation glyphosate-tolerant crops?

First-generation crops primarily relied on the modified EPSPS gene for tolerance. Second-generation crops incorporate both the modified EPSPS gene and Tra genes (like GOX) for enhanced and more robust glyphosate tolerance.

Topics Covered

BiotechnologyGeneticsHerbicide ResistanceGenetic EngineeringGene Function