UPSC MainsBOTANY-PAPER-II201215 Marks
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Q28.

How is light energy captured in green plants?

How to Approach

This question requires a detailed explanation of the photosynthetic process, focusing on how light energy is initially captured and then converted into chemical energy. The answer should cover the role of pigments like chlorophyll, the structure of chloroplasts, the light-dependent reactions, and the involvement of photosystems. A clear, step-by-step explanation is crucial, along with relevant biological terminology. The answer should demonstrate a strong understanding of plant physiology.

Model Answer

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Introduction

Photosynthesis, the cornerstone of life on Earth, is the process by which green plants and other organisms convert light energy into chemical energy. This process sustains nearly all life forms, directly or indirectly. The initial step in photosynthesis involves the capture of light energy by specialized pigments, primarily chlorophylls and carotenoids, housed within organelles called chloroplasts. Understanding how this capture occurs is fundamental to comprehending the entire photosynthetic pathway and its significance for global ecosystems. This answer will detail the mechanisms of light energy capture in green plants, outlining the key components and processes involved.

The Role of Pigments

Light energy is not directly usable by plants; it must first be absorbed by pigment molecules. These pigments are responsible for capturing specific wavelengths of light. The primary pigment involved is chlorophyll, which exists in several forms (chlorophyll a, chlorophyll b, etc.). Chlorophyll absorbs light most strongly in the blue and red portions of the electromagnetic spectrum, reflecting green light, which is why plants appear green.

  • Chlorophyll a: The primary photosynthetic pigment.
  • Chlorophyll b: An accessory pigment that broadens the range of light wavelengths that can be used for photosynthesis.
  • Carotenoids: Accessory pigments that absorb light in the blue-green region and protect chlorophyll from photo-oxidation. They also play a role in photoprotection.

Structure of Chloroplasts

Light capture occurs within chloroplasts, organelles found in plant cells. Chloroplasts have a complex internal structure:

  • Outer and Inner Membranes: These membranes enclose the chloroplast.
  • Stroma: The fluid-filled space inside the chloroplast, containing enzymes for the Calvin cycle.
  • Thylakoids: Flattened, sac-like membranes arranged in stacks called grana. The thylakoid membrane contains the chlorophyll and other pigments.
  • Lumen: The space inside the thylakoid.

Light-Dependent Reactions: Capturing Light Energy

The capture of light energy initiates the light-dependent reactions, which occur in the thylakoid membranes. These reactions involve two main photosystems: Photosystem II (PSII) and Photosystem I (PSI).

Photosystem II (PSII)

PSII absorbs light energy, exciting electrons in chlorophyll molecules. These energized electrons are passed along an electron transport chain (ETC). Water molecules are split (photolysis) to replace the lost electrons, releasing oxygen as a byproduct. This is the source of nearly all the oxygen in Earth’s atmosphere.

Photosystem I (PSI)

Electrons from the ETC eventually reach PSI, where they are re-energized by light absorption. These energized electrons are then used to reduce NADP+ to NADPH, a crucial reducing agent for the Calvin cycle.

Chemiosmosis and ATP Synthesis

As electrons move through the ETC, protons (H+) are pumped from the stroma into the thylakoid lumen, creating a proton gradient. This gradient drives the synthesis of ATP (adenosine triphosphate) through a process called chemiosmosis, catalyzed by the enzyme ATP synthase. This process is known as photophosphorylation.

Summary of Light Capture Process

Step Description Location
Light Absorption Pigments (chlorophyll, carotenoids) absorb light energy. Thylakoid membrane
Electron Excitation Light energy excites electrons in chlorophyll. Photosystems II & I
Water Splitting Water is split to replace electrons in PSII, releasing oxygen. Thylakoid lumen
Electron Transport Excited electrons move through the ETC. Thylakoid membrane
ATP Synthesis Proton gradient drives ATP synthesis via chemiosmosis. Thylakoid membrane
NADPH Formation Electrons reduce NADP+ to NADPH. Photosystem I

Conclusion

In conclusion, light energy capture in green plants is a complex process initiated by pigment molecules within chloroplasts. The light-dependent reactions, involving Photosystems II and I, convert light energy into chemical energy in the form of ATP and NADPH. This process is not only vital for plant survival but also sustains the vast majority of life on Earth by providing oxygen and the foundation for food chains. Further research into optimizing photosynthetic efficiency holds immense potential for addressing global food security and climate change challenges.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Photosystem
A complex of proteins and pigment molecules (including chlorophyll) that captures light energy and initiates the light-dependent reactions of photosynthesis.
Photolysis
The splitting of water molecules using light energy during the light-dependent reactions of photosynthesis, releasing oxygen, protons, and electrons.

Key Statistics

Approximately 100-170 billion tons of carbon are fixed by photosynthesis each year globally.

Source: Schlesinger, W. H. (1997). Biogeochemistry: An Analysis of Global Change. Academic Press.

Photosynthesis converts approximately 3-6% of the available solar energy into biomass.

Source: Raven, P. H., Evert, R. F., & Eichhorn, S. E. (2013). Biology of Plants (8th ed.). W. H. Freeman and Company.

Examples

C4 Plants

C4 plants, like corn and sugarcane, have evolved mechanisms to enhance light energy capture and minimize photorespiration, particularly in hot and dry environments. They utilize a different pathway for initial carbon fixation, resulting in higher photosynthetic rates.

Frequently Asked Questions

What happens to the energy that is not absorbed by chlorophyll?

The energy from wavelengths of light not absorbed by chlorophyll is either reflected or dissipated as heat. Accessory pigments help to capture some of these wavelengths, broadening the range of light used for photosynthesis.

Topics Covered

BiologyPlant SciencePhotosynthesisPlant PhysiologyLight Absorption