UPSC MainsZOOLOGY-PAPER-II201215 Marks
Q16.

Compare the structure and function of 30s and 70s ribosomes.

How to Approach

This question requires a comparative analysis of 30S and 70S ribosomes, focusing on their structural components and functional roles. The answer should begin by defining ribosomes and their importance in protein synthesis. A table comparing key structural features (size, composition, rRNA molecules, proteins) is crucial. Functional differences, particularly regarding their presence in different organisms and sensitivity to antibiotics, should also be highlighted. The answer should demonstrate a strong understanding of molecular biology concepts.

Model Answer

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Introduction

Ribosomes are complex molecular machines responsible for protein synthesis (translation) in all living organisms. They are not membrane-bound organelles and are found in both prokaryotic and eukaryotic cells. Ribosomes are classified into two main types based on their sedimentation coefficient during ultracentrifugation: 70S ribosomes found in prokaryotes, mitochondria, and chloroplasts, and 30S ribosomes which are a subunit of the 70S ribosome, and 80S ribosomes found in eukaryotic cytoplasm. These differences in structure and composition are crucial for understanding their function and are exploited by certain antibiotics to selectively inhibit bacterial protein synthesis. This answer will comprehensively compare the structure and function of 30S and 70S ribosomes.

Structural Comparison of 30S and 70S Ribosomes

The primary structural difference between 30S and 70S ribosomes lies in their size, composition, and the types of ribosomal RNA (rRNA) and proteins they contain. The 'S' value (Svedberg unit) is a measure of sedimentation rate and is related to size and shape.

Feature 30S Ribosome (Prokaryotic Subunit) 70S Ribosome (Prokaryotic Complete Ribosome)
Sedimentation Coefficient 30S 70S
Molecular Weight (Daltons) ~900,000 ~2,700,000
rRNA Molecules 16S rRNA 23S rRNA & 5S rRNA
Number of Proteins ~21 proteins ~55 proteins
Composition 16S rRNA + ~21 ribosomal proteins 30S subunit (16S rRNA + ~21 proteins) + 50S subunit (23S rRNA, 5S rRNA + ~34 proteins)
Location Part of the 70S ribosome in prokaryotes Prokaryotic cytoplasm, mitochondria, and chloroplasts

Functional Roles and Differences

Both 30S and 70S ribosomes perform the essential function of protein synthesis, but they differ in their specific roles and sensitivity to inhibitors.

30S Ribosomal Function

  • Initiation of Translation: The 30S subunit plays a crucial role in recognizing the Shine-Dalgarno sequence (in prokaryotes) or the Kozak sequence (in eukaryotes) on mRNA, which helps position the mRNA correctly for translation initiation.
  • tRNA Binding: It binds to the initiator tRNA carrying formylmethionine (in prokaryotes) or methionine (in eukaryotes) and ensures correct codon-anticodon pairing.
  • Decoding mRNA: The 30S subunit is primarily responsible for accurately decoding the mRNA sequence during translation.
  • Antibiotic Targets: The 30S subunit is a common target for several antibiotics, including streptomycin, tetracycline, and kanamycin. These antibiotics interfere with different stages of translation, such as initiation, elongation, or proofreading.

70S Ribosomal Function

  • Overall Protein Synthesis: The 70S ribosome, as a complete unit, carries out the entire process of protein synthesis – initiation, elongation, and termination.
  • Peptide Bond Formation: The 50S subunit (part of the 70S ribosome) contains the peptidyl transferase center, which catalyzes the formation of peptide bonds between amino acids.
  • Translocation: The 70S ribosome translocates along the mRNA, moving from one codon to the next.
  • Ribosome Recycling: After translation is complete, the 70S ribosome dissociates into its subunits for recycling.
  • Antibiotic Targets: The 70S ribosome is also a target for antibiotics like chloramphenicol and erythromycin, which primarily affect the 50S subunit.

Differences in Antibiotic Sensitivity

The structural differences between 30S and 70S ribosomes explain their differential sensitivity to antibiotics. Eukaryotic 80S ribosomes are less susceptible to antibiotics that target 70S ribosomes, providing a degree of selective toxicity. For example, tetracycline binds to the 30S subunit and prevents tRNA attachment, inhibiting bacterial protein synthesis without significantly affecting eukaryotic cells.

Evolutionary Significance

The presence of 70S ribosomes in mitochondria and chloroplasts supports the endosymbiotic theory, which proposes that these organelles originated from ancient bacteria. The similarity in ribosomal structure between these organelles and bacteria provides strong evidence for their bacterial ancestry.

Conclusion

In conclusion, 30S and 70S ribosomes, while both essential for protein synthesis, exhibit significant structural and functional differences. The 30S subunit is crucial for initiation and decoding, while the 70S ribosome represents the complete functional unit. These differences are exploited by antibiotics to selectively target bacterial protein synthesis and are also reflected in the evolutionary history of organelles like mitochondria and chloroplasts. Understanding these distinctions is fundamental to comprehending the intricacies of gene expression and developing effective antimicrobial strategies.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Sedimentation Coefficient (S)
The Svedberg unit (S) is a non-metric unit for measuring the sedimentation rate of particles in a fluid during centrifugation. It is a function of the particle's mass, shape, and density, and is used to characterize ribosomes and other macromolecules.
Shine-Dalgarno Sequence
A ribosomal binding site in prokaryotic mRNA that is located upstream of the start codon (AUG). It is complementary to a sequence on the 16S rRNA of the 30S ribosomal subunit, facilitating ribosome binding and translation initiation.

Key Statistics

Approximately 30% of a bacterial cell's dry weight is comprised of ribosomes.

Source: Lodish et al., Molecular Cell Biology, 4th edition (2000)

Antibiotic resistance is estimated to cause 700,000 deaths globally each year, and this number is projected to rise to 10 million by 2050 if no action is taken.

Source: Review on Antimicrobial Resistance (2016)

Examples

Tetracycline Resistance

Bacteria can develop resistance to tetracycline through various mechanisms, including the acquisition of genes encoding efflux pumps that actively transport the antibiotic out of the cell, or ribosomal protection proteins that alter the ribosome structure to prevent tetracycline binding.

Frequently Asked Questions

Why are antibiotics that target 70S ribosomes generally safe for humans?

Human cells utilize 80S ribosomes for protein synthesis. The structural differences between 70S and 80S ribosomes mean that antibiotics targeting 70S ribosomes have a much lower affinity for 80S ribosomes, minimizing their impact on human cells.

Topics Covered

BiologyMolecular BiologyRibosomesProtein SynthesisMolecular Biology