UPSC MainsAGRICULTURE-PAPER-II201312 Marks200 Words
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Q27.

Water potential of plant cells.

How to Approach

This question requires a clear understanding of water potential and its role in plant physiology. The approach should begin with defining water potential and its components. Then, explain how it operates across cell membranes and drives water movement. Illustrate with examples of different cell types and their water potential values. Finally, briefly discuss the significance of water potential in plant processes like nutrient uptake and transpiration. A structured approach with clear headings and subheadings is crucial for a comprehensive answer.

Model Answer

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Introduction

Water potential, a fundamental concept in plant physiology, dictates the direction of water movement within a plant and between the plant and its environment. It's essentially a measure of the free energy of water, expressed in Pascals (Pa) or megapascals (MPa). A more negative water potential indicates a greater tendency for water to move into that area. Understanding water potential is crucial for comprehending processes like nutrient absorption, transpiration, and maintaining turgor pressure, which is vital for plant growth and structural integrity. The concept was developed to explain the movement of water in plants, which is affected by both pressure and solute concentration.

Defining Water Potential

Water potential (Ψ) is defined as the potential energy of water per unit volume relative to pure water at atmospheric pressure and room temperature. It's a crucial factor determining the direction of water movement. Pure water has a water potential of zero. Adding solutes or applying pressure reduces the water potential, making it a negative value. It's a composite property influenced by two main components:

  • Solute Potential (Ψs): Always negative, it's the reduction in water potential due to the presence of dissolved solutes. Higher solute concentrations result in a more negative Ψs.
  • Pressure Potential (Ψp): Can be positive or negative. Positive pressure (turgor pressure) increases water potential, while negative pressure (tension) reduces it.

The equation relating these components is: Ψ = Ψs + Ψp

Water Potential Across Plant Cells

Water potential varies across different plant cells and tissues, driving water movement. Consider the following:

  • Root Cells: Have a more negative water potential than the surrounding soil due to the higher solute concentration needed for mineral absorption. This creates a gradient that draws water into the roots.
  • Xylem: The water potential in the xylem is very negative, especially in leaves undergoing transpiration. This negative pressure is crucial for the ascent of sap.
  • Mesophyll Cells: Have a more negative water potential than the air spaces within the leaf, facilitating water loss through transpiration.

Comparison of Water Potential Values (Example)

Cell Type Water Potential (MPa)
Pure Water 0
Root Cell (mineral absorption) -0.2 to -0.8
Xylem (at leaf surface) -1.0 to -3.0
Mesophyll Cell (during transpiration) -0.6 to -1.2

Osmosis and Water Movement

Water movement between cells is driven by osmosis – the movement of water across a semi-permeable membrane from a region of higher water potential to a region of lower water potential. The membrane allows water to pass but restricts the movement of solutes. For example, water moves from the root cortex cells (higher water potential) to the xylem (lower water potential).

Significance in Plant Processes

Water potential plays a crucial role in various plant processes:

  • Nutrient Uptake: Creates the driving force for water absorption and subsequent nutrient transport.
  • Transpiration: Maintains the tension column in the xylem, enabling water movement from roots to leaves.
  • Turgor Pressure: Maintains cell rigidity and supports plant structure. Wilting occurs when turgor pressure is lost due to low water potential.

Case Study: Drought Tolerance in Xerophytes

Case Study: Prosopis juliflora (Mesquite) This drought-tolerant plant exhibits several adaptations to maintain a more positive water potential during water scarcity. These include deep root systems to access groundwater, reduced leaf surface area to minimize transpiration, and the accumulation of osmolytes (e.g., proline) to lower solute potential and maintain cell turgor. Research by the ICAR-Central Arid Zone Research Institute (CAZRI) has focused on understanding these mechanisms to improve drought resilience in arid regions.

Conclusion

In conclusion, water potential is a critical determinant of water movement within plants, influencing processes from nutrient uptake to transpiration. Understanding the interplay of solute and pressure potentials, and their distribution across different cell types, is crucial for appreciating plant physiology. As climate change intensifies, comprehending and manipulating water potential mechanisms will be vital for ensuring food security and maintaining plant health, especially in drought-prone regions.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Osmosis
The movement of water across a semi-permeable membrane from a region of high water potential to a region of low water potential.
Turgor Pressure
The pressure exerted by the cell contents against the cell wall, a key component of water potential influencing cell rigidity and plant structure.

Key Statistics

The water potential in a typical plant cell can range from -0.2 MPa to -2.0 MPa, depending on the species and environmental conditions. (Source: Taiz and Zeiger, Plant Physiology, 2010)

Source: Taiz and Zeiger, Plant Physiology, 2010

Transpiration accounts for approximately 90% of the water absorbed by plants, demonstrating the critical role of water potential gradient in this process. (Source: Raven et al., Biology, 2017)

Source: Raven et al., Biology, 2017

Examples

Turgor Pressure and Wilting

When a plant lacks water, its cells lose turgor pressure, causing the plant to wilt. This is a direct consequence of a decrease in water potential within the cells.

Frequently Asked Questions

What is the difference between water potential and osmotic potential?

Water potential is the overall potential of water, considering both solute concentration and pressure. Osmotic potential is solely due to the solute concentration and is always negative.

Topics Covered

BiologyPlant PhysiologyWater RelationsOsmosisCell Physiology