UPSC MainsAGRICULTURE-PAPER-II201312 Marks200 Words
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Q23.

Mechanism of translocation of water from roots to leaves.

How to Approach

This question requires a detailed explanation of the mechanism of water translocation. The approach should be to first define transpiration and its role, then explain the cohesion-tension theory, focusing on the roles of xylem, root pressure, and the Guttation process. A structured response with clear headings and subheadings is crucial for clarity and completeness. Diagrams, though not possible here, would greatly enhance understanding. The answer should highlight the interplay of various forces involved in this process.

Model Answer

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Introduction

Water translocation, the movement of water from roots to leaves, is a vital process for plant survival, enabling photosynthesis and nutrient transport. This upward movement against gravity is a remarkable feat of nature, primarily driven by transpiration, the evaporation of water from aerial parts of the plant. The process is intricately linked to the plant's physiology and environmental factors. Understanding this mechanism is crucial for comprehending plant adaptation and agricultural practices. The cohesion-tension theory, proposed by Dixon and Holley, is the widely accepted explanation for this phenomenon.

The Role of Transpiration

Transpiration is the primary driver of water movement. It creates a negative pressure or tension in the leaves, which pulls water upwards. Factors influencing transpiration rates include humidity, temperature, wind speed, and light intensity. High transpiration rates increase the demand for water, accelerating the translocation process.

The Cohesion-Tension Theory

The cohesion-tension theory explains water movement based on three key properties of water: cohesion, adhesion, and surface tension.

  • Cohesion: Water molecules stick to each other due to hydrogen bonding. This creates a continuous column of water throughout the plant.
  • Adhesion: Water molecules also adhere to the walls of the xylem vessels, further supporting the water column against gravity.
  • Surface Tension: Reduces the likelihood of water column breakage.

The Xylem Pathway

Water moves through the plant via the xylem, a specialized vascular tissue. Xylem vessels are long, continuous tubes formed from dead cells, providing a low-resistance pathway for water transport. The tension created by transpiration pulls the water column upwards through the xylem.

Root Pressure and Guttation

While transpiration is the primary driver, root pressure also contributes, albeit to a lesser extent. Root pressure is the positive pressure that develops in the roots due to the active pumping of ions into the xylem. This pressure can force water upwards, especially at night when transpiration rates are low.

Guttation is the exudation of water droplets from leaves, often observed in the early morning. This phenomenon is a direct result of root pressure exceeding transpiration pull.

Summary of the Translocation Process

Step Process Role
1 Transpiration Creates tension/negative pressure in leaves
2 Cohesion Maintains continuous water column
3 Adhesion Prevents water column from collapsing
4 Xylem Transport Provides pathway for water movement
5 Root Pressure Contributes to water movement, especially at night

Impact of Environmental Factors

Environmental factors significantly influence water translocation. Drought conditions reduce water availability, limiting transpiration and impacting plant growth. High temperatures increase transpiration rates, potentially leading to water stress. Wind increases transpiration, while humidity reduces it.

Recent Research & Developments

Recent research focuses on understanding the role of aquaporins (water channel proteins) in regulating water transport within the plant. These proteins can facilitate or restrict water flow, allowing plants to fine-tune their water balance. Studies also investigate the impact of climate change on transpiration patterns and plant water use efficiency.

Conclusion

In conclusion, water translocation from roots to leaves is a complex process primarily driven by the cohesion-tension theory and transpiration. While root pressure plays a supporting role, the continuous water column facilitated by cohesion and adhesion, and the negative pressure generated by transpiration, are the key mechanisms. Understanding this process is vital for optimizing agricultural practices and ensuring plant health, especially in the face of changing climatic conditions. Further research into aquaporins promises to refine our understanding and potentially improve water use efficiency in crops.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Transpiration
The process by which water is lost from plants in the form of water vapor. It’s a critical process for water movement.
Cohesion-Tension Theory
The dominant theory explaining water transport in plants, based on the properties of water: cohesion, adhesion, and the creation of tension due to transpiration.

Key Statistics

A mature tree can transpire 200-300 liters of water per day.

Source: Knowledge Cutoff

Aquaporins can increase water permeability in plant membranes by up to 1000-fold.

Source: Knowledge Cutoff

Examples

Desert Plants

Desert plants like cacti have adapted to minimize water loss through reduced leaf surface area (spines) and specialized water storage tissues. Their transpiration rates are significantly lower than those of plants in wetter climates.

Rice Cultivation

In rice cultivation, guttation is a common phenomenon, especially in the early stages of growth, due to high root pressure and low transpiration rates.

Frequently Asked Questions

Why is water movement primarily upwards?

While water can move in any direction, the strong negative pressure generated by transpiration in the leaves creates a powerful driving force for upward movement. The cohesive and adhesive properties of water ensure that this force is transmitted all the way to the roots.

Can root pressure alone sustain water transport in a tall tree?

No. Root pressure alone is insufficient to overcome the gravitational force and move water to the top of tall trees. Transpiration-driven tension is the primary mechanism.

Topics Covered

BiologyPlant PhysiologyWater TransportXylemTranspiration