UPSC MainsZOOLOGY-PAPER-I201310 Marks150 Words
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Q5.

Describe hyoid apparatus in Frog or Necturus.

How to Approach

This question requires a detailed description of the hyoid apparatus in either a frog or Necturus (mudpuppy). A good answer will focus on the structure, components, functions, and evolutionary significance of the hyoid arch. It's crucial to illustrate with diagrams (though not possible here, mention it). The answer should be comparative, highlighting differences if possible, and emphasize the role of the hyoid apparatus in respiration and feeding. Structure the answer by first defining the hyoid apparatus, then detailing its components in the chosen animal, followed by its functions.

Model Answer

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Introduction

The hyoid apparatus is a series of bones located in the anterior neck, situated between the mandible and the larynx. It doesn’t articulate directly with any other bone, serving as a dynamic support for the tongue and larynx. In amphibians, the hyoid apparatus plays a crucial role in both respiration (buccal pumping) and feeding (tongue projection). While present in all vertebrates, its structure and function vary considerably. This answer will focus on the hyoid apparatus in the frog ( *Rana tigrina*), detailing its components and functional significance.

Hyoid Apparatus in Frog (*Rana tigrina*)

The hyoid apparatus in frogs is a complex structure composed of several cartilaginous elements. It’s significantly reduced compared to higher vertebrates, reflecting the amphibian’s unique mode of respiration and feeding.

Components of the Hyoid Apparatus

  • Hyoid Arch: The main supporting structure, forming a horseshoe-shaped arch.
  • Hyoid Body (Os Hyoideum): A single, centrally located cartilage.
  • Thyrohyoid: Connects the hyoid body to the thyroid cartilage of the larynx.
  • Ceratohyal: Forms the lower part of the hyoid arch, articulating with the operculum.
  • Branchial Arch: Though reduced, remnants of branchial arches are present, contributing to the support of the buccal cavity.
  • Glossohyal: Connects the hyoid body to the tongue, crucial for tongue projection.

These cartilaginous elements are connected by ligaments and muscles, allowing for considerable flexibility and movement.

Detailed Description of Components

The hyoid body is a relatively small, flattened cartilage. The ceratohyal arches extend posteriorly and laterally from the hyoid body. The glossohyal is a slender cartilage that projects anteriorly, providing attachment for the tongue muscles. The thyrohyoid connects the hyoid apparatus to the larynx, playing a role in vocalization and respiration. The arrangement of these cartilages allows for a wide range of movement, essential for capturing prey.

Functions of the Hyoid Apparatus in Frog

  • Buccal Pumping: The hyoid apparatus contributes to the lowering of the buccal cavity floor, creating negative pressure that draws air into the lungs.
  • Tongue Projection: The glossohyal and associated muscles enable the rapid and forceful projection of the tongue to capture insects. This is a key adaptation for insectivorous frogs.
  • Support for the Tongue and Larynx: Provides a skeletal framework for these structures.
  • Vocalization: The hyoid apparatus, along with the larynx, plays a role in producing frog calls.

The muscles attached to the hyoid apparatus are innervated by cranial nerves, allowing for precise control of its movements. The hyoid apparatus is crucial for the frog’s predatory lifestyle and its ability to thrive in diverse environments.

Comparison with Necturus

While a detailed comparison isn't the primary focus, it's worth noting that Necturus, being a fully aquatic salamander, possesses a more robust and complete hyoid apparatus compared to the frog. Necturus retains more of the branchial arches, reflecting its reliance on gill respiration throughout life. The hyoid apparatus in Necturus is primarily involved in buccal pumping for respiration and supporting the tongue for prey capture, similar to frogs, but with greater emphasis on aquatic adaptations.

Conclusion

In conclusion, the hyoid apparatus in frogs is a fascinating example of evolutionary adaptation. Its cartilaginous structure, composed of the hyoid body, ceratohyal, glossohyal, and thyrohyoid, is intricately linked to the frog’s unique respiratory and feeding mechanisms. The flexibility and muscular control of the hyoid apparatus enable efficient buccal pumping and rapid tongue projection, crucial for survival. Understanding the hyoid apparatus provides valuable insights into the amphibian’s physiology and evolutionary history.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Buccal Pumping
A method of respiration used by amphibians where air is forced into the lungs by movements of the buccal cavity floor.
Cartilage
A flexible connective tissue found in many areas of the body, including the joints, ears, nose, and the hyoid apparatus. It provides support and cushioning.

Key Statistics

Approximately 6,266 species of frogs have been described as of 2023 (AmphibiaWeb, University of California, Berkeley).

Source: AmphibiaWeb

Over 40% of amphibian species are currently threatened with extinction (IUCN Red List, 2023).

Source: IUCN Red List

Examples

Poison Dart Frogs

Poison dart frogs utilize their hyoid apparatus and associated muscles to produce distinctive calls for attracting mates and defending territory. The complexity of these calls varies between species.

Frequently Asked Questions

What is the evolutionary significance of the hyoid apparatus?

The hyoid apparatus evolved from the branchial arches of fish, becoming modified over time to support the tongue, larynx, and contribute to respiration in terrestrial vertebrates.

Topics Covered

ZoologyAmphibian PhysiologySkeletal SystemHyoid ApparatusAmphibians