UPSC MainsZOOLOGY-PAPER-I201325 Marks
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Q7.

Write an account of parental care in Amphibia.

How to Approach

This question requires a detailed understanding of the diverse parental care strategies exhibited by amphibians. The answer should cover the spectrum of care, from no care to extensive care, highlighting variations across different amphibian orders (Anura, Urodela, and Apoda). Focus on the mechanisms, evolutionary advantages, and ecological factors influencing parental care. Structure the answer by first defining parental care, then detailing it across different groups, and finally discussing the evolutionary significance.

Model Answer

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Introduction

Parental care, defined as any behavioral or physiological investment by a parent that increases the survival of its offspring, is a fascinating aspect of amphibian biology. While traditionally considered a group with minimal parental investment, amphibians exhibit a surprising diversity of care strategies, ranging from complete abandonment of eggs to elaborate nest building and prolonged larval care. This variation is influenced by factors like predation pressure, environmental conditions, and phylogenetic history. Understanding these strategies provides insights into amphibian reproductive ecology and evolutionary adaptations.

Parental Care in Amphibia: A Comprehensive Overview

Amphibian parental care is remarkably diverse, varying significantly across orders and even within families. It’s crucial to understand the context of their life cycle – many amphibians undergo metamorphosis, transitioning from aquatic larvae to terrestrial adults, which influences the type and duration of care.

1. Anura (Frogs and Toads)

Frogs generally exhibit less parental care compared to salamanders. However, significant variations exist:

  • No Care: Most anurans simply deposit eggs in water and leave, relying on yolk reserves for larval development.
  • Nest Building: Some species construct foam nests (e.g., Leptodactylus pentadactylus) to protect eggs from desiccation and predation. These nests are often built near water and guarded by the male.
  • Egg Guarding: Males of certain species (e.g., Dendrobates poison dart frogs) guard the eggs, keeping them moist and removing predators.
  • Transporting Tadpoles: A remarkable example is the marsupial frog, Gastrotheca, where the female carries developing tadpoles on her back until metamorphosis.
  • Paternal Care of Tadpoles: Pipa pipa (Surinam toad) embeds eggs in its back, and the skin grows over them, providing protection and nourishment until fully formed froglets emerge.

2. Urodela (Salamanders)

Salamanders generally exhibit more extensive parental care than frogs, particularly in terrestrial species.

  • Egg Guarding: Many salamanders guard their eggs, protecting them from predators and fungal infections.
  • Nest Building: Some species build nests under rocks or logs, providing a humid environment for egg development.
  • Ventilation of Eggs: Salamanders like Ambystoma maculatum (spotted salamander) fan their eggs with their tails to increase oxygenation.
  • Parental Feeding: In some species, parents may provide unfertilized eggs as food for developing larvae.
  • Direct Development: Certain salamanders (e.g., Bolitoglossa) exhibit direct development, bypassing the larval stage. Females provide nourishment to the developing embryos within the eggs, resulting in miniature adults hatching.

3. Apoda (Caecilians)

Caecilians, being burrowing amphibians, exhibit unique parental care strategies.

  • Oophagy: Some caecilians exhibit oophagy, where developing embryos feed on unfertilized eggs within the mother’s oviduct.
  • Dermatophagy: In some species, the young feed on the mother’s skin, which is shed and enriched with nutrients.
  • Viviparity: Certain caecilians are viviparous, giving birth to live young.

Evolutionary Significance of Parental Care

The evolution of parental care in amphibians is linked to several factors:

  • Predation Pressure: Higher predation rates often favor increased parental care to enhance offspring survival.
  • Environmental Conditions: Harsh or unpredictable environments may necessitate parental protection and provisioning.
  • Life History Traits: Species with larger eggs and slower development often exhibit more parental care.
  • Phylogenetic Constraints: Evolutionary history plays a role in shaping the types of care observed in different groups.
Order Typical Parental Care Examples
Anura Egg deposition, limited guarding, foam nests, transport Dendrobates, Leptodactylus pentadactylus, Gastrotheca
Urodela Egg guarding, nest building, ventilation, direct development Ambystoma maculatum, Bolitoglossa
Apoda Oophagy, dermatophagy, viviparity Various caecilian species

Conclusion

Parental care in amphibians is a diverse and fascinating phenomenon, shaped by a complex interplay of ecological pressures and evolutionary history. While many species exhibit minimal care, others demonstrate remarkable adaptations to protect and nourish their offspring. This variation highlights the plasticity of amphibian reproductive strategies and their sensitivity to environmental changes. Further research is crucial to understand the long-term consequences of habitat loss and climate change on amphibian parental care and population viability.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Oophagy
A form of parental nutrition in which developing embryos feed on unfertilized eggs.
Viviparity
Giving birth to live young that have developed inside the mother’s body.

Key Statistics

Approximately 40% of amphibian species exhibit some form of parental care (Duellman, 1999).

Source: Duellman, W. E. (1999). *Cusco amphibians*. University of Kansas Natural History Museum.

Salamanders exhibit parental care in approximately 60% of species (Wells, 2007).

Source: Wells, K. D. (2007). *The ecology and behavior of amphibians*. University of Chicago Press.

Examples

Poison Dart Frogs

Male poison dart frogs (<em>Dendrobates</em>) carry tadpoles on their backs to small pools of water, often in bromeliads, providing them with food (unfertilized eggs) and protection.

Frequently Asked Questions

Why do some frogs abandon their eggs while others provide extensive care?

The level of parental care is often correlated with predation risk and environmental conditions. Species in more dangerous environments or with limited resources tend to exhibit more care to increase offspring survival.

Topics Covered

ZoologyAnimal BehaviorParental CareAmphibiansReproduction