UPSC MainsBOTANY-PAPER-I201412 Marks
Q23.

T.S. of cyathium of Euphorbia.

How to Approach

This question requires a detailed, diagrammatic description of the transverse section (T.S.) of a cyathium, specifically in *Euphorbia*. The answer should focus on identifying and explaining the various floral parts present in the cyathium, highlighting its unique structure as a specialized inflorescence. A clear, labelled diagram is crucial. The answer should also explain the evolutionary significance of this structure. Focus on the arrangement of structures like involucres, cyathophylls, glands, and the reduced female flower.

Model Answer

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Introduction

The cyathium is a highly specialized inflorescence characteristic of the genus *Euphorbia* (family Euphorbiaceae). It represents a significant evolutionary adaptation, often mistaken for a single flower due to its compact structure. The cyathium is a pseudanthium – a structure that appears to be a single flower but is actually a cluster of reduced flowers surrounded by modified bracts. Understanding its anatomy is crucial for comprehending the reproductive strategies and phylogenetic relationships within the Euphorbiaceae family. This detailed description will focus on the T.S. of a typical *Euphorbia* cyathium, highlighting its key features.

T.S. of Cyathium of *Euphorbia*

The transverse section of a *Euphorbia* cyathium reveals a complex arrangement of structures. The following points detail the key features observed in a typical T.S.:

1. Involucre

The outermost layer is formed by the involucre, which is a whorl of bracts (involucral bracts). These bracts are often green and leaf-like, providing protection to the developing floral structures. They are arranged in multiple rows and may be slightly fused at their bases.

2. Cyathophylls

Inside the involucre are cyathophylls, which are modified leaves that surround the flowers. These are typically brightly colored (often yellow or red) and serve to attract pollinators. They are arranged in a spiral or whorled pattern.

3. Glands (Nectaries)

A prominent feature of the cyathium is the presence of numerous glands, often located at the base of the cyathophylls. These glands secrete nectar, attracting insects for pollination. They appear as circular or oval structures in the T.S.

4. Male Flowers

The male flowers are highly reduced, typically consisting of a single stamen (filament and anther). They are numerous and arranged around the central female flower. The anthers are often small and dehisce explosively, releasing pollen.

5. Female Flower

The female flower is also reduced, consisting of a single pistil (stigma, style, and ovary). The ovary is usually superior and contains one or more ovules. The stigma is often branched or lobed to increase the surface area for pollen reception.

6. Peduncle

The cyathium is borne on a peduncle, which is a stalk that connects the inflorescence to the main stem. The vascular bundles are visible within the peduncle.

Here's a tabular representation summarizing the key components:

Component Description Function
Involucre Whorl of bracts Protection of floral parts
Cyathophylls Modified leaves Attract pollinators
Glands Nectar-secreting structures Attract pollinators
Male Flower Single stamen Pollen production
Female Flower Single pistil Ovule protection and fertilization

7. Vascular Bundles

The cyathium exhibits a complex vascular arrangement. Vascular bundles are present in the peduncle, involucre, and cyathophylls, providing nourishment and support to the developing floral structures. The arrangement of vascular bundles can be used to identify the different parts of the cyathium.

Pollination Mechanism: The cyathium relies on insect pollination, primarily by small beetles and flies. The nectar glands attract these insects, which inadvertently transfer pollen from the male flowers to the stigma of the female flower. The explosive pollen release mechanism further aids in pollination.

Conclusion

The T.S. of the *Euphorbia* cyathium reveals a remarkable adaptation for reproduction. Its unique structure, characterized by reduced floral parts and specialized bracts and glands, demonstrates the evolutionary pressures that have shaped plant morphology. Understanding the anatomy of the cyathium is essential for appreciating the diversity and complexity of plant reproductive strategies. Further research into the genetic mechanisms controlling cyathium development could provide valuable insights into plant evolution and breeding.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Pseudanthium
A pseudanthium is an inflorescence that resembles a single flower, but is actually a cluster of many small flowers tightly packed together with modified bracts. The cyathium of *Euphorbia* is a classic example.
Involucre
A whorl of bracts surrounding a flower or inflorescence, often providing protection to the developing floral structures. In the cyathium, the involucre is a key identifying feature.

Key Statistics

The genus *Euphorbia* comprises over 2,000 species, making it one of the largest genera of flowering plants.

Source: Christenhusz, M. J. M., & Byng, J. W. (2016). The number of known plant species in the world and its distribution. *Phytokeys*, *54*, 201–211.

Approximately 75% of *Euphorbia* species are succulents, adapted to arid and semi-arid environments.

Source: Data based on knowledge cutoff - botanical databases and taxonomic literature (as of late 2023).

Examples

Poinsettia (*Euphorbia pulcherrima*)

Poinsettia, a popular ornamental plant, exhibits a brightly colored cyathium, with the cyathophylls (modified leaves) being the prominent red structures often mistaken for petals. This adaptation enhances its attractiveness to pollinators.

Frequently Asked Questions

What is the evolutionary advantage of the cyathium structure?

The cyathium structure likely evolved to enhance pollination efficiency. The compact arrangement of flowers and the presence of nectar glands attract pollinators, while the explosive pollen release mechanism ensures effective pollen transfer. It also provides protection to the developing female flower.

Topics Covered

BotanyPlant BiologyPlant AnatomyInflorescenceEuphorbia