UPSC MainsAGRICULTURE-PAPER-I201512 Marks
Q14.

What is photoperiodism? Write at least one example of shortday plant, longday plant and day neutral plant.

How to Approach

This question requires a clear understanding of photoperiodism, its mechanisms, and examples. The approach should begin by defining photoperiodism and explaining its biological significance. Then, illustrate the concept with examples of short-day, long-day, and day-neutral plants, highlighting their flowering responses to varying day lengths. A structured response with clear headings and concise explanations will be crucial for a comprehensive answer. Finally, enrichment items will provide further context and detail.

Model Answer

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Introduction

Photoperiodism is a vital phenomenon in the plant kingdom, influencing various developmental stages, most notably flowering. It refers to the physiological response of plants to the relative lengths of day and night. This response governs a wide array of processes, including seed germination, stem growth, and leaf abscission. Understanding photoperiodism is crucial for agricultural practices, enabling farmers to manipulate flowering times and optimize crop yields, especially in regions with varying day lengths throughout the year. The discovery of photoperiodism by Garner and Allard in 1920, while studying soybeans, laid the foundation for modern understanding of plant responses to light duration.

What is Photoperiodism?

Photoperiodism is the ability of plants to sense and respond to the length of daylight hours. This response is mediated by phytochrome, a photoreceptor pigment that exists in two interconvertible forms: Pr (inactive) and Pfr (active). The ratio of red (R) to far-red (FR) light determines the equilibrium between these forms. During daylight, red light converts Pr to Pfr, while during night, Pfr reverts to Pr. The duration of the dark period is critical in determining the plant's response.

Types of Plants Based on Photoperiodic Response

Plants are broadly classified into three categories based on their photoperiodic responses:

Short-Day Plants (SDPs)

SDPs flower when the day length is shorter than a critical threshold, or when the night length exceeds a specific duration. They are often referred to as "long-night plants" because flowering is triggered by extended darkness.

  • Mechanism: In SDPs, the phytochrome system senses the length of the dark period. A long night period leads to a high concentration of Pr, which inhibits flowering.
  • Example: Chrysanthemums (Chrysanthemum morifolium) are classic examples of SDPs. They require nights of 12-14 hours to flower. Commercial chrysanthemum production often involves artificially shortening day length to induce flowering out of season.
  • Other Examples: Poinsettias, Strawberries, Rice

Long-Day Plants (LDPs)

LDPs flower when the day length is longer than a critical threshold, or when the night length is shorter than a specific duration. They are often called "short-night plants" because flowering is triggered by a brief period of darkness.

  • Mechanism: In LDPs, a short night period allows for the accumulation of Pfr, which promotes flowering. A long night period reverts Pfr to Pr, inhibiting flowering.
  • Example: Spinach (Spinacia oleracea) is a well-known LDP. It requires days of 14-16 hours to flower. Farmers in regions with shorter daylight hours often use artificial lighting to extend the day length and induce flowering.
  • Other Examples: Lettuce, Radish, Barley

Day-Neutral Plants (DNPs)

DNPs flower irrespective of day length. Their flowering is primarily influenced by other factors like maturity, temperature, and moisture.

  • Mechanism: DNPs do not have a photoperiodic response; their flowering is not directly controlled by day length.
  • Example: Tomatoes (Solanum lycopersicum) are a common example of DNPs. Their flowering is influenced more by the age of the plant and environmental conditions than by the length of the day.
  • Other Examples: Corn, Cucumbers, Sunflowers
Plant Type Critical Day Length Night Length Requirement Example
Short-Day Plant Shorter than critical threshold Longer than critical duration Chrysanthemum
Long-Day Plant Longer than critical threshold Shorter than critical duration Spinach
Day-Neutral Plant No effect No effect Tomato

Applications and Significance

Understanding photoperiodism has significant implications for agriculture:

  • Crop Improvement: Plant breeders can manipulate photoperiodic genes to develop varieties that flower at specific times of the year, aligning with desired harvest schedules.
  • Greenhouse Cultivation: Artificial lighting and shading can be used to control day length in greenhouses, allowing for year-round production of crops that are normally seasonal.
  • Ornamental Horticulture: Photoperiodism is exploited to control flowering in ornamental plants, ensuring blooms at desired times for commercial sales.

The Indian Council of Agricultural Research (ICAR) actively researches photoperiodism in various crops to improve productivity and adapt to changing climatic conditions. Recent research focuses on understanding the genetic mechanisms underlying photoperiodic responses in rice and other staple crops.

Conclusion

In conclusion, photoperiodism is a crucial physiological mechanism that governs flowering and other developmental processes in plants. The classification of plants into short-day, long-day, and day-neutral categories based on their photoperiodic responses highlights the complexity of this phenomenon. Understanding and manipulating photoperiodism has significant implications for agriculture, allowing for increased crop yields, improved quality, and adaptability to diverse environmental conditions. Continued research in this area is essential for ensuring food security and sustainable agricultural practices in a changing world.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Phytochrome
A photoreceptor protein in plants that mediates the plant’s response to red and far-red light, playing a crucial role in photoperiodism and other light-regulated developmental processes.
Critical Day Length
The specific day length that triggers flowering in long-day plants or inhibits flowering in short-day plants. This value varies among plant species and cultivars.

Key Statistics

Global greenhouse vegetable production is estimated to be over 300 million tonnes annually, with photoperiod manipulation being a key factor in optimizing yields. (Source: FAOSTAT, 2021)

Source: FAOSTAT

Studies show that manipulating photoperiod in rice varieties can increase grain yield by up to 15-20% in certain regions. (Source: ICAR Research Report, 2018 – Knowledge Cutoff)

Source: ICAR Research Report

Examples

Artificial Day Length Control in Chrysanthemum Production

Commercial chrysanthemum growers often use black-out curtains or other light-blocking methods to create artificial short days, inducing flowering even during periods of long natural days. This allows for year-round production and market availability.

Frequently Asked Questions

Why are some plants considered "long-night plants" even though they are called short-day plants?

The terminology can be confusing. “Short-day plants” flower when the *day* is short, which means the *night* is long. Thus, they are more accurately described as long-night plants.

Topics Covered

BiologyBotanyPlant PhysiologyPlant HormonesFloweringPhotoperiodism