UPSC MainsAGRICULTURE-PAPER-II201512 Marks
Q28.

Name the process by which carbohydrates are synthesized in plants. Narrate its mechanism.

How to Approach

This question requires a detailed explanation of photosynthesis, the process by which plants synthesize carbohydrates. The approach should begin by defining photosynthesis and its significance. Then, the mechanism should be broken down into light-dependent and light-independent reactions (Calvin cycle), detailing each step. Diagrams can be mentally visualized to aid understanding. Finally, factors affecting photosynthesis should be briefly mentioned. The answer should be structured around these key components to ensure a comprehensive and well-organized response.

Model Answer

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Introduction

Photosynthesis, derived from the Greek words "photo" (light) and "synthesis" (putting together), is the cornerstone of life on Earth, serving as the primary mechanism for converting light energy into chemical energy. It's the process by which plants, algae, and certain bacteria use sunlight, water, and carbon dioxide to create oxygen and energy in the form of sugars. This process directly supports almost all ecosystems, providing the foundation for food chains and maintaining atmospheric oxygen levels. Recent advancements in understanding photosynthetic efficiency, like research into C4 and CAM plants, are crucial for addressing food security challenges in a changing climate.

Photosynthesis: An Overview

Photosynthesis can be broadly divided into two stages: the light-dependent reactions and the light-independent reactions (Calvin Cycle). Both are intricately linked and occur within chloroplasts, specifically in the thylakoid membranes (light-dependent) and the stroma (light-independent).

Light-Dependent Reactions (Photochemical Phase)

These reactions occur in the thylakoid membranes of the chloroplasts and require light energy. The process involves several key steps:

  • Light Absorption: Chlorophyll and other pigment molecules (carotenoids, phycobilins) within photosystems I (PSI) and II (PSII) absorb light energy.
  • Photosystem II (PSII): Light energy excites electrons in PSII, which are then passed along an electron transport chain (ETC). Water molecules are split (photolysis) to replace these electrons, releasing oxygen as a byproduct. The equation for photolysis is: 2H2O → 4H+ + 4e- + O2
  • Electron Transport Chain (ETC): As electrons move down the ETC, energy is released, which is used to pump protons (H+) from the stroma into the thylakoid lumen, creating a proton gradient.
  • Photosystem I (PSI): Electrons reach PSI, where they are re-energized by light. These energized electrons are then used to reduce NADP+ to NADPH.
  • ATP Synthesis (Chemiosmosis): The proton gradient created by the ETC drives the synthesis of ATP from ADP and inorganic phosphate through a process called chemiosmosis, facilitated by ATP synthase. This process is also known as photophosphorylation.

The net result of the light-dependent reactions is the production of ATP, NADPH, and oxygen.

Light-Independent Reactions (Calvin Cycle / Carbon Fixation)

These reactions occur in the stroma of the chloroplast and do not directly require light, but rely on the products of the light-dependent reactions (ATP and NADPH). The Calvin cycle can be divided into three phases:

  • Carbon Fixation: Carbon dioxide from the atmosphere is combined with ribulose-1,5-bisphosphate (RuBP), a five-carbon molecule. This reaction is catalyzed by the enzyme RuBisCO (Ribulose-1,5-bisphosphate carboxylase/oxygenase).
  • Reduction: The resulting six-carbon molecule is unstable and immediately breaks down into two molecules of 3-phosphoglycerate (3-PGA). ATP and NADPH are used to convert 3-PGA into glyceraldehyde-3-phosphate (G3P).
  • Regeneration: Some G3P molecules are used to synthesize glucose and other carbohydrates. The remaining G3P molecules are used to regenerate RuBP, allowing the cycle to continue.

The overall equation for photosynthesis is:

6CO2 + 6H2O + Light Energy → C6H12O6 + 6O2

Factors Affecting Photosynthesis

Several factors influence the rate of photosynthesis, including:

  • Light Intensity: Photosynthesis increases with light intensity up to a certain point.
  • Carbon Dioxide Concentration: Higher CO2 concentrations generally increase the rate of photosynthesis, up to a limit.
  • Temperature: Photosynthesis has an optimal temperature range. Extremely high or low temperatures can inhibit enzyme activity.
  • Water Availability: Water stress can lead to stomatal closure, reducing CO2 uptake and inhibiting photosynthesis.
  • Nutrient Availability: Nutrients like nitrogen and magnesium are essential for chlorophyll synthesis and enzyme function.

C4 and CAM Photosynthesis

Some plants have evolved alternative photosynthetic pathways, C4 and CAM, to overcome limitations in hot, dry environments. C4 plants (e.g., maize, sugarcane) spatially separate carbon fixation and the Calvin cycle. CAM plants (e.g., cacti, succulents) temporally separate these processes.

Feature C4 Photosynthesis CAM Photosynthesis
Spatial Separation Yes - Carbon fixation in mesophyll cells, Calvin cycle in bundle sheath cells No - Both processes occur in the same cells
Temporal Separation No Yes - Carbon fixation at night, Calvin cycle during the day
RuBisCO Efficiency Higher - Reduced photorespiration Higher - Reduced photorespiration

Conclusion

In conclusion, photosynthesis is a vital biochemical process enabling plants to convert light energy into chemical energy in the form of carbohydrates. It involves intricate light-dependent and light-independent reactions, each with specific steps and requirements. Understanding this process and its influencing factors is critical for improving crop yields, addressing food security, and mitigating the impacts of climate change. Further research into optimizing photosynthetic efficiency, particularly through genetic engineering, holds significant promise for sustainable agriculture.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Photolysis
The splitting of water molecules using light energy during photosynthesis, releasing electrons, protons, and oxygen.
RuBisCO
Ribulose-1,5-bisphosphate carboxylase/oxygenase, the enzyme responsible for carbon fixation in the Calvin cycle. It's the most abundant enzyme on Earth.

Key Statistics

Approximately 150 billion metric tons of carbon are fixed by photosynthesis annually worldwide. (Source: NASA Earth Observatory, knowledge cutoff)

Source: NASA Earth Observatory

RuBisCO has a relatively low catalytic efficiency, with a turnover number (number of molecules converted per minute) of only 2-4. (Source: Biochemistry textbooks, knowledge cutoff)

Source: Biochemistry Textbooks

Examples

Maize (Corn)

Maize is a classic example of a C4 plant, demonstrating efficient photosynthesis in warm climates with high light intensity.

Cactus

Cacti are a prime example of CAM plants, adapted to arid environments by opening their stomata at night to absorb carbon dioxide and performing photosynthesis during the day.

Frequently Asked Questions

What is photorespiration, and why is it a problem?

Photorespiration occurs when RuBisCO binds with oxygen instead of carbon dioxide, reducing photosynthetic efficiency. It's more prevalent in C3 plants under hot, dry conditions.

How do C4 and CAM plants improve upon the standard C3 photosynthesis?

C4 and CAM plants minimize photorespiration by concentrating CO2 around RuBisCO, allowing for more efficient carbon fixation and higher photosynthetic rates, especially in challenging environments.

Topics Covered

BiologyBotanyPlant PhysiologyPhotosynthesisCarbon FixationPlant Metabolism