UPSC MainsBOTANY-PAPER-I201515 Marks
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Q9.

Draw well labelled diagram of the T.S. of Teleutosorus of Puccinia and give a brief description of the same.

How to Approach

This question requires a detailed understanding of plant pathology, specifically the reproductive structure of *Puccinia* species. The answer should focus on accurately depicting the T.S. of the teleutosorus with clear labeling and a concise description of its components and function. A well-labeled diagram is crucial, and the description should highlight the key features and their significance in the life cycle of the rust fungus. The answer should demonstrate knowledge of fungal morphology and plant-pathogen interactions.

Model Answer

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Introduction

Rust fungi, belonging to the order Uredinales, are obligate plant pathogens causing significant economic losses in agriculture. *Puccinia graminis*, the black stem rust of wheat, is a notorious example. The teleutosorus is a crucial structure in the life cycle of these fungi, representing the overwintering stage and facilitating sexual reproduction. It is a specialized sorus containing teleutospores, dikaryotic cells that undergo meiosis to produce basidiospores. Understanding the morphology of the teleutosorus is fundamental to comprehending the disease cycle and developing effective control strategies.

T.S. of Teleutosorus of *Puccinia* – A Detailed Description

The teleutosorus is a dark-colored, often elongated or cylindrical structure formed on the host plant, typically towards the end of the growing season. A transverse section (T.S.) reveals a complex arrangement of cells. The following describes the key features observed in a T.S. of a mature teleutosorus:

1. Structure and Layers

The teleutosorus is embedded within the host tissue and consists of several layers:

  • Peridium: The outermost protective layer, formed from host cells, providing a barrier against environmental stresses.
  • Teleutospores: The main component, consisting of numerous, dark-colored, thick-walled, binucleate (dikaryotic) cells. These are the overwintering spores.
  • Sterigmata: Hyphal branches that support the teleutospores. They arise from the underlying fungal tissue.
  • Subhypodermis: A layer of host cells beneath the peridium.
  • Mesophyll: The underlying host tissue providing nutrients.

2. Teleutospores – Morphology and Function

Teleutospores are typically dark brown or black due to the presence of melanin in their thick walls. They are generally oval or spherical in shape and possess a prominent pore (germ pore) at one end. This pore facilitates the emergence of the basidium during germination.

3. Basidium Formation and Basidiospores

Upon exposure to favorable conditions (moisture and temperature), the teleutospore germinates, forming a four-celled basidium. Meiosis occurs within the basidium, resulting in the formation of four haploid basidiospores. These basidiospores are forcibly discharged and dispersed by wind, initiating the next stage of the fungal life cycle.

4. Diagram of T.S. of Teleutosorus

T.S. of Teleutosorus of Puccinia

Key: 1. Peridium, 2. Teleutospores, 3. Sterigmata, 4. Subhypodermis, 5. Mesophyll

5. Significance in Disease Cycle

The teleutosorus plays a critical role in the completion of the *Puccinia* life cycle. It allows the fungus to survive unfavorable conditions (winter) and undergo sexual reproduction, generating genetic diversity through meiosis. The resulting basidiospores infect alternate hosts, continuing the disease cycle.

Conclusion

The teleutosorus of *Puccinia* is a complex structure essential for the survival and reproduction of rust fungi. Its detailed morphology, including the peridium, teleutospores, and sterigmata, reflects its function in overwintering and initiating the sexual phase of the life cycle. Understanding this structure is crucial for developing effective disease management strategies aimed at disrupting the fungal life cycle and minimizing crop losses. Further research into the genetic mechanisms governing teleospore formation and basidiospore production could lead to novel approaches for rust disease control.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Teleutospores
Dikaryotic, thick-walled, overwintering spores of rust fungi, produced within the teleutosorus. They undergo meiosis to form basidiospores.
Sorus (plural: Sori)
A cluster of spores produced by fungi, often visible as a powdery or colored mass on the surface of a host plant. Different types of sori are characteristic of different fungal species and life cycle stages.

Key Statistics

Wheat rust diseases, including black stem rust caused by *Puccinia graminis*, can cause yield losses of up to 70% in susceptible wheat varieties.

Source: FAO (Food and Agriculture Organization of the United Nations), 2023 (Knowledge Cutoff)

Global annual losses due to wheat rust diseases are estimated to be over $3 billion.

Source: Global Rust Initiative, 2022 (Knowledge Cutoff)

Examples

Ug99 Strain of *Puccinia graminis*

The Ug99 strain of black stem rust, first identified in Uganda in 1999, is a particularly virulent race that threatens global wheat production. It overcame many rust-resistant genes previously deployed in wheat varieties, necessitating the development of new resistance strategies.

Frequently Asked Questions

What is the difference between aecia, uredinia, and teleutosori?

Aecia, uredinia, and teleutosori are different types of sori produced by rust fungi at different stages of their life cycle. Aecia produce sperm cells, uredinia produce urediniospores (for asexual reproduction during the growing season), and teleutosori produce teleutospores (for overwintering and sexual reproduction).

Topics Covered

BotanyPlant PathologyFungal BiologyPlant DiseasesMicroscopy