UPSC MainsZOOLOGY-PAPER-I201515 Marks
Q23.

What is chromosome painting? Comment on its principles and scope.

How to Approach

This question requires a detailed understanding of chromosome painting, a molecular cytogenetic technique. The answer should begin with a clear definition of the technique, followed by a discussion of its underlying principles – primarily fluorescence in situ hybridization (FISH). The scope section should cover its applications in various fields like karyotyping, cancer research, and evolutionary biology. Structuring the answer with subheadings like 'Principles', 'Procedure', and 'Applications' will enhance clarity. Mentioning specific probes and their origins will demonstrate depth of knowledge.

Model Answer

0 min read

Introduction

Chromosome painting is a fluorescence in situ hybridization (FISH) technique used to visualize entire chromosomes or large chromosomal regions. Developed in the early 1990s, it revolutionized cytogenetics by providing a powerful tool for identifying chromosomal abnormalities and studying genome organization. Unlike traditional karyotyping which relies on banding patterns, chromosome painting utilizes fluorescently labeled DNA probes that bind to specific chromosome regions, allowing for clear visualization and identification even in complex karyotypes. This technique has become indispensable in both clinical diagnostics and fundamental research.

Principles of Chromosome Painting

The core principle behind chromosome painting is Fluorescence In Situ Hybridization (FISH). This involves using DNA probes, which are single-stranded DNA fragments complementary to specific sequences on chromosomes. These probes are labeled with fluorescent dyes (fluorochromes), allowing them to be visualized under a fluorescence microscope.

  • Probe Preparation: Probes are typically generated by flow sorting of chromosomes, followed by PCR amplification of the isolated DNA. This ensures that the probe contains DNA representative of the entire chromosome or a specific region.
  • Hybridization: The fluorescently labeled probes are denatured (separated into single strands) and hybridized to metaphase chromosomes that have also been denatured.
  • Visualization: After hybridization, the chromosomes are washed to remove unbound probes, and then visualized using a fluorescence microscope. Each chromosome or region appears brightly colored, corresponding to the fluorochrome used to label the probe.

Procedure of Chromosome Painting

The process of chromosome painting involves several key steps:

  1. Sample Preparation: Typically, cells in metaphase are used, as the chromosomes are most condensed and easily visualized.
  2. Chromosome Preparation: Cells are treated with a hypotonic solution to swell the chromosomes, followed by fixation in methanol/acetic acid.
  3. Probe Denaturation & Hybridization: Probes are denatured by heating and then applied to the chromosomes. Hybridization is carried out under specific conditions of temperature and salt concentration.
  4. Washing: Excess probes are removed by washing in a stringent buffer.
  5. Counterstaining & Visualization: Chromosomes are counterstained with DAPI (4′,6-diamidino-2-phenylindole) to visualize all DNA, and then observed under a fluorescence microscope.

Scope and Applications of Chromosome Painting

Chromosome painting has a wide range of applications in various fields:

  • Karyotyping and Chromosomal Aberration Detection: It allows for rapid and accurate identification of chromosomal abnormalities such as translocations, deletions, and inversions, even in complex karyotypes where banding patterns are difficult to interpret.
  • Cancer Research: Chromosome painting is crucial in identifying chromosomal rearrangements that are characteristic of specific cancers. For example, the Philadelphia chromosome (a translocation between chromosomes 9 and 22) can be easily identified using chromosome painting.
  • Prenatal Diagnosis: It can be used to detect chromosomal abnormalities in fetal cells obtained through amniocentesis or chorionic villus sampling.
  • Evolutionary Biology: Chromosome painting helps in understanding the evolutionary relationships between species by comparing their chromosome organization and identifying conserved synteny (the order of genes on chromosomes).
  • Genome Mapping: It aids in mapping genes to specific chromosomes and identifying regions of homology between different genomes.

Types of Probes Used

Different types of probes are used depending on the application:

Probe Type Target Application
Whole Chromosome Probes Entire chromosome Karyotyping, detection of whole chromosome aneuploidy
Centromeric Probes Centromeric region Detection of centromere abnormalities
Telomeric Probes Telomeric regions Detection of telomere shortening or abnormalities
Region-Specific Probes Specific chromosomal region Mapping genes, detecting microdeletions/duplications

Conclusion

Chromosome painting has emerged as a powerful and versatile technique in cytogenetics, offering significant advantages over traditional karyotyping. Its ability to visualize entire chromosomes or specific regions with high resolution has revolutionized the diagnosis of chromosomal abnormalities and advanced our understanding of genome organization and evolution. Continued advancements in probe technology and imaging techniques promise to further expand the scope and applications of chromosome painting in the future, particularly in personalized medicine and genomic research.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Synteny
The conserved order of genes on chromosomes across different species, indicating evolutionary relationships.

Key Statistics

The global FISH market was valued at USD 1.6 billion in 2023 and is projected to reach USD 2.5 billion by 2032, growing at a CAGR of 5.1% from 2024 to 2032.

Source: Global Market Insights, 2024 (Knowledge Cutoff: April 2024)

FISH testing is performed on approximately 500,000 prenatal samples annually in the United States alone.

Source: American Journal of Human Genetics, 2022 (Knowledge Cutoff: April 2024)

Examples

Chronic Myeloid Leukemia (CML)

Chromosome painting is routinely used to detect the Philadelphia chromosome, a hallmark of CML, which results from a reciprocal translocation between chromosomes 9 and 22. This translocation creates the BCR-ABL fusion gene, driving uncontrolled cell proliferation.

Frequently Asked Questions

What are the limitations of chromosome painting?

Chromosome painting requires metaphase spreads, which can be difficult to obtain from some tissues. It also relies on the availability of high-quality probes and can be affected by probe specificity and hybridization conditions. Furthermore, it cannot detect subtle chromosomal rearrangements like inversions within a single chromosome arm.

Topics Covered

Science & TechnologyBiologyGeneticsCytogeneticsChromosome AnalysisMolecular Biology