UPSC MainsAGRICULTURE-PAPER-II201612 Marks
Q15.

Give an account of mechanism of protein synthesis. Compare it with prokaryotes and eukaryotes.

How to Approach

This question requires a detailed explanation of protein synthesis, a fundamental biological process. The approach should begin with defining protein synthesis and outlining its stages (transcription and translation). Then, a comparison between prokaryotic and eukaryotic protein synthesis mechanisms must be presented, highlighting differences in mRNA processing, ribosome structure, and initiation factors. Finally, a concise conclusion summarizing the key distinctions is essential. The answer should be structured logically, using clear headings and bullet points for enhanced readability.

Model Answer

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Introduction

Protein synthesis, the cornerstone of cellular life, is the process by which cells build proteins. These proteins are vital for virtually every cellular function, from catalyzing biochemical reactions to transporting molecules and providing structural support. The process involves two main stages: transcription, where DNA is copied into messenger RNA (mRNA), and translation, where mRNA is decoded to assemble amino acids into a polypeptide chain. Recent advancements in CRISPR-Cas9 technology have allowed scientists to manipulate protein synthesis pathways, demonstrating its critical role in genetic engineering and disease treatment. Understanding the nuances of this process, especially the differences between prokaryotes and eukaryotes, is crucial for comprehending the diversity of life.

Mechanism of Protein Synthesis: A Detailed Overview

Protein synthesis is a complex process occurring in two main stages: transcription and translation. Both stages involve intricate molecular machinery and are crucial for cellular function.

Transcription

Transcription is the process of creating an RNA copy from a DNA template. This occurs in two steps:

  • Initiation: RNA polymerase binds to the promoter region on the DNA.
  • Elongation: RNA polymerase moves along the DNA template, synthesizing a complementary mRNA molecule.
  • Termination: RNA polymerase reaches a termination signal, releasing the mRNA molecule.

Translation

Translation is the process of converting the mRNA sequence into a polypeptide chain. It occurs on ribosomes, which are composed of ribosomal RNA (rRNA) and proteins. The process involves three stages:

  • Initiation: The ribosome binds to the mRNA and a transfer RNA (tRNA) molecule carrying the first amino acid (usually methionine).
  • Elongation: tRNA molecules, each carrying a specific amino acid, sequentially bind to the mRNA codon, and the amino acids are linked together by peptide bonds.
  • Termination: The ribosome reaches a stop codon on the mRNA, releasing the polypeptide chain.

Comparison of Protein Synthesis in Prokaryotes and Eukaryotes

While the fundamental principles of protein synthesis are similar in prokaryotes and eukaryotes, there are significant differences in the details of the process.

Feature Prokaryotes Eukaryotes
Location Cytoplasm Nucleus (Transcription), Cytoplasm (Translation)
mRNA Processing Minimal; mRNA is directly used for translation. Extensive; includes 5' capping, splicing (removal of introns), and 3' polyadenylation.
Ribosomes 70S (50S + 30S subunits) 80S (60S + 40S subunits)
Initiation Factors Fewer initiation factors (IF1, IF2, IF3) More complex initiation factors (eIFs) - e.g., eIF4E, eIF2, eIF3. eIF4E binds to the 5' cap, crucial for scanning the mRNA.
Coupling of Transcription and Translation Coupled; translation begins while transcription is still in progress. Uncoupled; transcription and translation occur separately.
Introns Rarely present Commonly present; require splicing

The differences arise from the complexity of eukaryotic cells, which have a nucleus and other membrane-bound organelles. The nuclear membrane necessitates mRNA processing steps that are absent in prokaryotes, which lack a nucleus. The larger ribosomal subunits and more complex initiation factors in eukaryotes reflect the greater complexity of their protein synthesis machinery.

Examples of Differences in Action

Consider the synthesis of actin, a crucial protein for eukaryotic cell structure and movement. The actin mRNA undergoes extensive processing, including splicing to remove introns. This is absent in prokaryotic systems. Similarly, the initiation of actin translation requires multiple eukaryotic initiation factors (eIFs), a process absent in bacteria.

Case Study: Antibiotic Resistance in Bacteria

Case Study Title: Impact of Antibiotics on Bacterial Protein Synthesis

Description: Many antibiotics target bacterial protein synthesis machinery. For instance, tetracycline inhibits bacterial protein synthesis by binding to the 30S ribosomal subunit, preventing tRNA from binding. This highlights the differences between bacterial (70S) and eukaryotic (80S) ribosomes, which allows for selective targeting. The emergence of antibiotic resistance often involves mutations in ribosomal proteins or the acquisition of genes encoding proteins that modify antibiotics, thereby circumventing their inhibitory effects.

Outcome: Understanding the differences in protein synthesis pathways between bacteria and eukaryotes is crucial for developing new antibiotics that are effective against resistant strains.

FAQ

FAQ Question: What is the role of tRNA in protein synthesis?

FAQ Answer: tRNA molecules act as adaptors, bringing the correct amino acid to the ribosome based on the mRNA codon sequence. Each tRNA has an anticodon that complements a specific mRNA codon.

Scheme

Scheme Name: Biotechnology Parks Scheme (DBT, India)

Description: This scheme, under the Department of Biotechnology (DBT), aims to promote research and development in biotechnology, including protein engineering and synthetic biology, which often relies on understanding and manipulating protein synthesis pathways. It provides infrastructure and funding for research institutions and biotech companies.

Year: 2002

Conclusion

In conclusion, protein synthesis is a fundamental biological process vital for all life forms. While the basic mechanism remains consistent, significant differences exist between prokaryotic and eukaryotic protein synthesis, primarily due to the structural and organizational differences between these cell types. These differences offer opportunities for targeted therapeutic interventions, as exemplified by antibiotic development. Future research focusing on manipulating protein synthesis pathways holds immense potential for advancements in medicine and biotechnology.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Codon
A three-nucleotide sequence in mRNA that codes for a specific amino acid or a stop signal during translation.
Ribosome
A complex molecular machine responsible for protein synthesis in all living cells. It reads mRNA and links amino acids to form a polypeptide chain.

Key Statistics

Approximately 20,000-25,000 genes are estimated to be present in the human genome, each potentially encoding a different protein.

Source: NCBI Knowledge Base

The average rate of translation in mammalian cells is approximately 10-20 amino acids per second.

Source: Based on knowledge cutoff - rates can vary significantly.

Examples

Insulin Synthesis

The synthesis of insulin in pancreatic beta cells showcases the complexity of eukaryotic protein synthesis, involving extensive mRNA processing and the action of numerous regulatory factors.

Frequently Asked Questions

Why is mRNA processing more complex in eukaryotes?

Eukaryotic mRNA processing is more complex due to the presence of introns in genes, which must be removed by splicing. This ensures that only the coding regions of the gene are translated into protein.

Topics Covered

BiologyBiochemistryGeneticsCentral dogma, genetic code, ribosomes, mRNA, tRNA