UPSC MainsBOTANY-PAPER-I201615 Marks
Q30.

Heart wood and Sap wood

How to Approach

This question requires a comparative analysis of heartwood and sapwood, two crucial components of secondary xylem. The answer should define both terms, detail their formation, structural differences, functions, and economic importance. A tabular comparison will be highly effective. Focus on anatomical features and physiological roles. The answer should demonstrate a strong understanding of plant anatomy and its relevance to forestry and timber production.

Model Answer

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Introduction

Secondary growth in plants leads to the formation of wood, a complex tissue providing structural support and facilitating transport. Within wood, two distinct regions are identifiable: heartwood and sapwood. These regions differ significantly in their composition, function, and appearance. Heartwood, the central, older wood, provides strength and durability, while sapwood, the outer, younger wood, is responsible for water and mineral transport. Understanding the characteristics of both is crucial for appreciating wood properties and sustainable forest management.

Heartwood and Sapwood: A Detailed Comparison

Both heartwood and sapwood are components of secondary xylem, formed by the vascular cambium. However, their development and functions diverge significantly.

1. Formation and Development

Sapwood is the initially formed secondary xylem, actively involved in water conduction. As the xylem cells age and become non-functional in water transport, they undergo a series of biochemical changes, collectively known as heartwood formation. These changes involve the deposition of substances like tannins, resins, gums, and oils into the cell lumens. This process makes the wood darker, harder, and more resistant to decay.

2. Anatomical Differences

The anatomical differences between heartwood and sapwood are key to their distinct properties. Sapwood cells generally have thinner walls and are more permeable due to the presence of open pits. Heartwood cells, on the other hand, have thicker walls and often contain tyloses (balloon-like ingrowths of parenchyma cells into xylem vessels) and resin ducts, reducing permeability.

Feature Sapwood Heartwood
Color Light colored Dark colored (brown, reddish)
Cell Wall Thickness Thinner Thicker
Permeability Highly permeable Less permeable
Cell Contents Starch, tannins (less) Tannins, resins, oils, gums (more)
Weight Lighter Heavier
Susceptibility to Decay More susceptible Less susceptible
Function Water and mineral transport Structural support, durability

3. Functional Differences

Sapwood’s primary function is the efficient conduction of water and minerals from the roots to the leaves. This is facilitated by the open structure of its vessels. Heartwood, having lost its conductive capacity, primarily provides mechanical strength and support to the plant. The deposited substances also protect the wood from fungal and insect attacks.

4. Economic Importance

Both heartwood and sapwood have economic value, but for different reasons. Sapwood is easier to work with and is often used in applications where permeability is desired, such as pulpwood for paper production. Heartwood, due to its durability and resistance to decay, is highly valued for construction, furniture making, and other applications requiring long-lasting wood. The proportion of heartwood to sapwood in a tree influences its timber quality and market value.

5. Species Variation

The extent of heartwood formation varies significantly among different tree species. Some species, like teak (Tectona grandis), naturally produce a large proportion of heartwood, making them highly prized for their durability. Others, like poplar (Populus spp.), have a relatively small amount of heartwood, and their wood is more susceptible to decay.

Conclusion

In conclusion, heartwood and sapwood represent distinct stages in the development and function of secondary xylem. Sapwood facilitates vital transport processes, while heartwood provides structural integrity and protection. Understanding the differences between these two wood types is essential for effective forest management, timber utilization, and appreciating the complex physiology of plants. The proportion and quality of heartwood are key determinants of timber value, influencing sustainable forestry practices and economic returns.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Secondary Xylem
The vascular tissue produced by the vascular cambium, responsible for water and mineral transport, and providing structural support to the plant. It forms the bulk of wood.

Key Statistics

Globally, approximately 3.4 billion cubic meters of wood are harvested annually (FAO, 2020 - knowledge cutoff). A significant portion of this harvest targets heartwood due to its superior properties.

Source: Food and Agriculture Organization of the United Nations (FAO)

Approximately 50% of the dry weight of heartwood can be comprised of extractives like tannins and resins (based on knowledge cutoff 2023).

Source: Forest Products Laboratory, USDA

Examples

Teak Wood

Teak (<em>Tectona grandis</em>) is renowned for its high heartwood content, which is exceptionally durable and resistant to decay, making it ideal for shipbuilding, outdoor furniture, and construction in tropical climates.

Frequently Asked Questions

What causes heartwood to be darker in color?

The darker color of heartwood is primarily due to the deposition of secondary metabolites like tannins, resins, and other phenolic compounds within the cell lumens. These compounds contribute to the wood's durability and resistance to decay.

Topics Covered

BotanyPlant AnatomyWood anatomyXylemPlant physiology