UPSC MainsZOOLOGY-PAPER-II201615 Marks
Q8.

Explain the salient features in the 'theory of natural selection'. Add a special comment on 'synthetic theory of evolution'.

How to Approach

This question requires a detailed understanding of Darwin’s theory of natural selection and its subsequent refinement into the modern synthetic theory. The answer should begin by outlining the core tenets of natural selection – variation, inheritance, selection, and time. Then, it should explain how the synthetic theory integrated Mendelian genetics with Darwinian evolution, addressing the gaps in the original theory. A comparative approach highlighting the differences and advancements will be beneficial. Structure the answer into an introduction, a detailed body covering both theories, and a concise conclusion.

Model Answer

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Introduction

The theory of evolution, a cornerstone of modern biology, explains the diversity of life on Earth. While various evolutionary theories existed prior, Charles Darwin’s ‘On the Origin of Species’ (1859) revolutionized the field with his theory of natural selection. This theory proposed a mechanism for evolutionary change based on observable phenomena. However, Darwin lacked a complete understanding of the source of variation and inheritance. The subsequent integration of Mendelian genetics with Darwinian evolution in the 20th century led to the ‘synthetic theory of evolution’, providing a more comprehensive and robust explanation of evolutionary processes. This answer will detail the salient features of natural selection and then elaborate on the synthetic theory, highlighting its advancements.

Darwin’s Theory of Natural Selection

Darwin’s theory of natural selection is based on four key observations:

  • Variation: Individuals within a population exhibit variations in their traits. These variations can be morphological, physiological, or behavioral.
  • Inheritance: Traits are passed down from parents to offspring. Darwin, however, didn’t understand the mechanism of inheritance.
  • Selection: Due to limited resources, there is a struggle for existence. Individuals with traits that provide an advantage in a particular environment are more likely to survive and reproduce. This is ‘natural selection’.
  • Time: Over long periods, these advantageous traits become more common in the population, leading to adaptation and, eventually, the formation of new species.

Darwin’s theory explained adaptation as a result of gradual changes accumulating over generations. He used the example of finches in the Galapagos Islands, demonstrating how beak shapes varied based on available food sources. He also proposed the concept of ‘descent with modification’, suggesting that all life forms share a common ancestor.

Limitations of Darwin’s Theory

Despite its groundbreaking nature, Darwin’s theory had limitations:

  • Lack of understanding of inheritance: Darwin proposed ‘pangenesis’, a flawed theory of inheritance. He couldn’t explain how traits were consistently passed down.
  • Origin of variation: Darwin couldn’t explain the source of variation. He simply observed that it existed.
  • Slow rate of evolution: Some evolutionary changes seemed to occur too rapidly to be explained solely by natural selection.

The Synthetic Theory of Evolution

The synthetic theory of evolution, developed between the 1930s and 1950s, integrated Darwinian natural selection with Mendelian genetics, resolving many of the limitations of the original theory. Key contributors included Ronald Fisher, J.B.S. Haldane, Sewall Wright, Theodosius Dobzhansky, and Ernst Mayr.

The core tenets of the synthetic theory are:

  • Genetic Variation: Variation arises from random mutations in genes and recombination during sexual reproduction.
  • Gene Pool: Evolution acts on the gene pool of a population, not on individuals.
  • Hardy-Weinberg Equilibrium: This principle describes the conditions under which allele and genotype frequencies remain constant in a population. Deviations from this equilibrium indicate that evolution is occurring.
  • Mechanisms of Evolutionary Change: The synthetic theory identifies five mechanisms of evolutionary change:
    • Mutation: The ultimate source of new genetic variation.
    • Gene Flow: The transfer of genes between populations.
    • Genetic Drift: Random fluctuations in allele frequencies, particularly in small populations.
    • Non-random mating: Mate choice based on specific traits.
    • Natural Selection: Differential survival and reproduction based on heritable traits.
  • Speciation: The formation of new species occurs through reproductive isolation, preventing gene flow between populations.

Comparison of Darwin’s and Synthetic Theory

Feature Darwin’s Theory Synthetic Theory
Source of Variation Unexplained Random mutations and recombination
Mechanism of Inheritance Pangenesis (incorrect) Mendelian genetics
Unit of Evolution Individual Population (gene pool)
Focus Adaptation Evolutionary forces and mechanisms

Conclusion

In conclusion, Darwin’s theory of natural selection laid the foundation for our understanding of evolution, but it lacked a complete explanation for the origin of variation and its inheritance. The synthetic theory of evolution successfully integrated Mendelian genetics with Darwinian principles, providing a more comprehensive and robust framework for understanding the evolutionary process. This synthesis continues to be refined with advancements in molecular biology and genomics, offering deeper insights into the mechanisms driving the diversity of life on Earth. The ongoing research in fields like epigenetics and horizontal gene transfer further expands our understanding of evolutionary processes.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Adaptation
The process by which organisms become better suited to their environment through the evolution of specific traits.
Gene Flow
The transfer of genetic material from one population to another, often through migration and interbreeding.

Key Statistics

Approximately 99% of all species that have ever lived on Earth are now extinct (based on fossil records as of 2023).

Source: The Paleobiology Database

Humans share approximately 98% of their DNA with chimpanzees (Source: National Human Genome Research Institute, knowledge cutoff 2023).

Source: National Human Genome Research Institute

Examples

Industrial Melanism in Peppered Moths

During the Industrial Revolution in England, the frequency of dark-colored peppered moths increased due to pollution darkening tree bark. This provided camouflage against predators, demonstrating natural selection in action.

Frequently Asked Questions

Is evolution still happening today?

Yes, evolution is an ongoing process. We observe it in the development of antibiotic resistance in bacteria, pesticide resistance in insects, and the evolution of viruses.

Topics Covered

BiologyEvolutionNatural SelectionEvolutionary TheorySynthetic Theory