UPSC MainsZOOLOGY-PAPER-I201715 Marks
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Q7.

Give the structure of cnidoblast, the defensive organ in Cnidaria. Also, describe its mechanism of action.

How to Approach

This question requires a detailed understanding of the cnidoblast, a unique feature of Cnidarians. The answer should begin with a clear definition of cnidoblasts and their function. Then, a step-by-step description of the cnidoblast's structure, including its key components like the nematocyst, operculum, and cnidocil, is crucial. Finally, the mechanism of action – from stimulation to nematocyst discharge and its effects – needs to be explained comprehensively. Diagrams, though not possible here, would be highly beneficial in an exam setting.

Model Answer

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Introduction

Cnidarians, including jellyfish, corals, and hydra, are characterized by specialized stinging cells called cnidocytes, containing organelles called cnidoblasts. These cnidoblasts are responsible for capturing prey, defense against predators, and attachment to substrates. The cnidoblast houses a complex intracellular structure called the nematocyst, a harpoon-like organelle capable of delivering venom. Understanding the structure and mechanism of action of cnidoblasts is fundamental to comprehending the biology and ecological role of Cnidarians. This answer will detail the structure of the cnidoblast and elucidate the intricate process of nematocyst discharge.

Structure of the Cnidoblast

The cnidoblast is a specialized cell containing the nematocyst. Its structure can be divided into several key components:

  • Cnidocil: A sensory hair-like structure protruding from the cell. It acts as a trigger for nematocyst discharge.
  • Operculum: A lid-like structure covering the nematocyst’s opening. It prevents premature discharge.
  • Nematocyst: The capsule containing the coiled, thread-like tubule. This is the actual stinging apparatus.
  • Cytoplasm: The general cellular contents surrounding the nematocyst.
  • Nucleus: The control center of the cell.

The nematocyst itself is a complex structure. It consists of:

  • Capsule: The outer covering of the nematocyst, containing the coiled tubule.
  • Tubule: A hollow, thread-like structure that is rapidly everted during discharge. It can be barbed, sticky, or venomous.
  • Volvent filament: A spirally coiled filament within the capsule, contributing to the force of eversion.

Mechanism of Action

The discharge of the nematocyst is one of the fastest biological processes known, occurring within microseconds. The mechanism involves the following steps:

  1. Stimulation: The cnidocil is stimulated by mechanical contact (e.g., prey touching the tentacle) or chemical signals.
  2. Ion Flux: Stimulation causes an influx of calcium ions (Ca2+) into the cnidoblast cytoplasm.
  3. Osmotic Pressure Change: The increased Ca2+ concentration leads to a rapid increase in osmotic pressure within the nematocyst capsule.
  4. Operculum Rupture: The increased pressure causes the operculum to rupture, exposing the nematocyst’s opening.
  5. Tubule Eversion: The coiled tubule is rapidly everted, propelled by the high osmotic pressure and the volvent filament. This eversion is incredibly forceful.
  6. Penetration & Injection: The everted tubule penetrates the prey’s skin or surface. Depending on the type of nematocyst, it may inject venom, entangle the prey, or adhere to the surface.
  7. Anchoring & Contraction: After penetration, the tubule anchors itself, and the capsule contracts, pulling the prey closer to the cnidarian’s mouth.

Types of Nematocysts

There are several types of nematocysts, each adapted for a specific function:

Nematocyst Type Function Characteristics
Penetrants Prey capture and defense Barbed tubules that pierce the prey’s skin and inject venom.
Glutinants Attachment Small, sticky tubules that adhere to surfaces.
Volvents Prey entanglement Thread-like tubules that wrap around the prey.

It’s important to note that once a nematocyst is discharged, it is not reusable. The cnidoblast must synthesize a new nematocyst to regain its stinging capability.

Conclusion

In conclusion, the cnidoblast is a remarkable example of cellular specialization, enabling Cnidarians to thrive in diverse marine environments. Its intricate structure, particularly the nematocyst, and the rapid, forceful mechanism of its discharge are crucial for prey capture, defense, and locomotion. Understanding these features provides valuable insight into the evolutionary success and ecological role of this fascinating phylum. Further research into the molecular mechanisms governing nematocyst discharge could have implications for biomedical applications, such as developing novel drug delivery systems.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Cnidocyte
A specialized stinging cell in Cnidarians that contains the cnidoblast and nematocyst. It is the functional unit responsible for stinging.

Key Statistics

The discharge of a nematocyst can occur in as little as 700 nanoseconds, making it one of the fastest biological processes known.

Source: Schmidt, G. R. (1979). Biology of nematocysts. Annual Review of Physiology, 41(1), 379-405.

Approximately 75% of cnidarian species are marine, highlighting their dominance in oceanic ecosystems.

Source: Barnes, R. S. K., Calow, P., Olive, P. J. W., Golding, D. W., & Spicer, J. I. (1987). The invertebrates: a synthesis. Blackwell Scientific Publications.

Examples

Portuguese Man-of-War

The Portuguese Man-of-War (Physalia physalis) is a siphonophore, a colonial organism related to jellyfish, famous for its extremely potent nematocysts. Contact with its tentacles can cause severe pain, welts, and even systemic symptoms in humans.

Frequently Asked Questions

Are all Cnidarians equally venomous?

No, the potency of the venom and the type of nematocysts vary significantly among different Cnidarian species. Some species, like the box jellyfish (Chironex fleckeri), are highly venomous and potentially lethal to humans, while others have relatively mild stings.

Topics Covered

ZoologyInvertebrate ZoologyCnidariaCell BiologyDefense Mechanisms