UPSC MainsZOOLOGY-PAPER-I201710 Marks150 Words
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Q15.

Describe 'FISH: Principle, Procedure and Applications'.

How to Approach

This question requires a detailed explanation of Fluorescence In Situ Hybridization (FISH). The answer should cover the principle behind the technique (how it works), the procedure involved in performing FISH, and its diverse applications in various fields, particularly in genetics and diagnostics. Structure the answer by first defining FISH, then detailing the procedural steps, and finally listing its applications with specific examples. Focus on clarity and conciseness, adhering to the word limit.

Model Answer

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Introduction

Fluorescence In Situ Hybridization (FISH) is a powerful cytogenetic technique used to detect and localize the presence or absence of specific DNA sequences on chromosomes. Developed in the 1980s, FISH revolutionized genetic diagnostics by allowing visualization of DNA sequences directly within cells, offering a significant advantage over traditional karyotyping. It utilizes fluorescent probes that bind to complementary DNA sequences, enabling the identification of chromosomal abnormalities, gene copy number variations, and even specific gene mutations. This technique has become indispensable in clinical genetics, cancer research, and prenatal diagnostics.

Principle of FISH

The fundamental principle of FISH relies on the hybridization of a labeled DNA probe to its complementary sequence on a target chromosome or DNA sequence. The probe, a short single-stranded DNA fragment, is labeled with a fluorescent dye (fluorophore). When the probe encounters its complementary sequence, it binds (hybridizes) through base pairing (A with T, and G with C). This hybridization is then visualized using a fluorescence microscope, where the fluorophore emits light upon excitation, revealing the location of the target sequence.

Procedure of FISH

The FISH procedure involves several key steps:

  • Probe Preparation: DNA probes are designed to target specific genomic regions. These probes are labeled with fluorophores like FITC, Texas Red, or Cy5, each emitting a different color.
  • Sample Preparation: Cells or tissues are fixed (e.g., with formaldehyde) to preserve their morphology. Chromosomes are often denatured (separated into single strands) by heating.
  • Hybridization: The denatured probe is added to the denatured sample. Hybridization occurs under controlled conditions (temperature, salt concentration) to allow specific binding.
  • Washing: Unbound probe is removed by washing.
  • Detection & Visualization: The sample is counterstained with a DNA dye (e.g., DAPI) to visualize all chromosomes. A fluorescence microscope is used to detect the fluorescent signal from the hybridized probe.

Applications of FISH

FISH has a wide range of applications across various fields:

  • Chromosomal Aberration Detection: Identifying aneuploidies (abnormal chromosome number, e.g., Down syndrome – Trisomy 21), deletions, duplications, and translocations.
  • Cancer Diagnostics: Detecting gene amplifications (e.g., HER2/neu in breast cancer) and chromosomal rearrangements associated with specific cancers.
  • Prenatal Diagnosis: Rapidly detecting chromosomal abnormalities in fetal cells obtained through amniocentesis or chorionic villus sampling.
  • Gene Mapping: Locating specific genes on chromosomes.
  • Species Identification: Identifying different species based on their unique DNA sequences.
  • Microbial Detection: Identifying specific bacteria or viruses in clinical samples.

Comparative Genomic Hybridization (CGH) using FISH (FISH-CGH): A variation of FISH used to detect copy number variations across the entire genome. It compares the genomic DNA of a test sample to a normal reference sample.

Application Specific Example
Cancer Diagnostics Detecting the Philadelphia chromosome (translocation between chromosomes 9 and 22) in Chronic Myeloid Leukemia (CML).
Prenatal Diagnosis Identifying Trisomy 18 (Edwards syndrome) in a fetus.
Gene Mapping Confirming the location of the BRCA1 gene on chromosome 17.

Conclusion

FISH remains a cornerstone of modern cytogenetics and molecular diagnostics. Its ability to visualize specific DNA sequences directly within cells provides invaluable information for diagnosing genetic disorders, understanding cancer development, and advancing our knowledge of genome organization. Continued advancements in probe technology and imaging techniques are expanding the applications of FISH, promising even more precise and efficient genetic analysis in the future. The technique’s versatility ensures its continued relevance in both research and clinical settings.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Fluorophore
A fluorescent chemical compound that can re-emit light upon absorbing light of a shorter wavelength. Commonly used labels in FISH.
Hybridization
The process of forming a double-stranded nucleic acid molecule (DNA or RNA) by complementary base pairing between two single-stranded nucleic acid molecules.

Key Statistics

The global FISH market was valued at USD 1.4 billion in 2023 and is projected to reach USD 2.2 billion by 2032, growing at a CAGR of 5.2% from 2024 to 2032.

Source: Global Market Insights, 2024 (Knowledge Cutoff: April 2024)

FISH testing is performed on approximately 70% of newly diagnosed breast cancer cases to assess HER2 status.

Source: American Cancer Society, 2023 (Knowledge Cutoff: April 2024)

Examples

Detection of HER2 Amplification

In breast cancer, FISH is routinely used to determine if the HER2 gene is amplified. HER2 amplification indicates that the cancer cells produce too much of the HER2 protein, making them more aggressive. This information guides treatment decisions, as patients with HER2-positive breast cancer benefit from targeted therapies like trastuzumab (Herceptin).

Frequently Asked Questions

What is the difference between FISH and karyotyping?

Karyotyping provides an overall view of chromosome number and structure, but it has limited resolution. FISH can detect smaller deletions or duplications and can be targeted to specific DNA sequences, offering higher resolution and specificity.

Topics Covered

ZoologyMolecular BiologyGeneticsMolecular TechniquesCytogenetics