UPSC MainsZOOLOGY-PAPER-II201720 Marks
Q20.

Give a comprehensive account of gaseous transport in blood.

How to Approach

This question requires a detailed understanding of the mechanisms involved in gaseous transport, specifically oxygen and carbon dioxide, within the blood. The answer should cover the roles of hemoglobin, plasma, and the physiological processes of loading, unloading, and regulation. A structured approach, detailing the transport of each gas separately, followed by a discussion of factors influencing transport, is recommended. Focus on the biochemical and physiological aspects, and avoid overly simplistic explanations.

Model Answer

0 min read

Introduction

Gaseous transport is a vital physiological process ensuring oxygen delivery to tissues and removal of carbon dioxide, a metabolic waste product. This exchange is crucial for cellular respiration and maintaining homeostasis. The blood serves as the primary medium for this transport, utilizing both dissolved gases and specialized carrier molecules. Efficient gaseous transport is dependent on the interplay between the lungs, blood, and tissues, and is regulated by various physiological mechanisms. Understanding this process is fundamental to comprehending respiratory physiology and related clinical conditions.

Oxygen Transport

Oxygen transport in blood is primarily mediated by hemoglobin, a protein found in red blood cells (erythrocytes). A small amount of oxygen (approximately 1.5%) is dissolved directly in plasma, but this contributes minimally to overall oxygen delivery.

Hemoglobin and Oxygen Binding

  • Hemoglobin Structure: Hemoglobin consists of four globin chains (two alpha and two beta in adults), each containing a heme group with an iron atom. Each iron atom can bind one oxygen molecule.
  • Oxygen-Hemoglobin Dissociation Curve: This curve illustrates the relationship between partial pressure of oxygen (PO2) and hemoglobin saturation. It is sigmoidal, indicating cooperative binding – the binding of one oxygen molecule increases the affinity for subsequent oxygen molecules.
  • Factors Affecting Oxygen Binding: Several factors influence the oxygen-hemoglobin dissociation curve:
    • pH (Bohr Effect): Decreased pH (increased acidity) reduces hemoglobin's affinity for oxygen, promoting oxygen release in tissues.
    • Temperature: Increased temperature decreases hemoglobin's affinity for oxygen.
    • PCO2: Increased PCO2 decreases hemoglobin's affinity for oxygen (Haldane effect).
    • 2,3-Bisphosphoglycerate (2,3-BPG): Increased levels of 2,3-BPG decrease hemoglobin's affinity for oxygen, facilitating oxygen release.

Carbon Dioxide Transport

Carbon dioxide transport in blood is more complex than oxygen transport, utilizing multiple mechanisms. Approximately 7-10% of CO2 is dissolved in plasma, 20-25% is bound to hemoglobin (carbaminohemoglobin), and the majority (60-70%) is transported as bicarbonate ions (HCO3-).

Mechanisms of CO2 Transport

  • Dissolved CO2: CO2 dissolves in plasma, contributing to a small portion of total CO2 transport.
  • Carbaminohemoglobin: CO2 binds to the amino groups of hemoglobin, forming carbaminohemoglobin. This binding is influenced by PO2 – higher PO2 promotes CO2 release (Haldane effect).
  • Bicarbonate Formation: The majority of CO2 is converted to bicarbonate ions (HCO3-) within red blood cells through the following reaction, catalyzed by carbonic anhydrase:

    CO2 + H2O ⇌ H2CO3 ⇌ H+ + HCO3-

    The bicarbonate ions are then transported out of the red blood cells into the plasma in exchange for chloride ions (chloride shift). This maintains electrical neutrality.

Regulation of Gaseous Transport

Gaseous transport is tightly regulated by several mechanisms to ensure adequate oxygen delivery and CO2 removal.

  • Ventilation-Perfusion Matching: The lungs regulate ventilation (airflow) and perfusion (blood flow) to match oxygen supply with metabolic demand.
  • Chemoreceptors: Central chemoreceptors in the medulla oblongata and peripheral chemoreceptors in the carotid and aortic bodies detect changes in PO2, PCO2, and pH, triggering adjustments in ventilation rate and depth.
  • Erythropoiesis: The production of red blood cells is regulated by erythropoietin, a hormone released by the kidneys in response to hypoxia (low oxygen levels).
Gas Primary Transport Mechanism Secondary Mechanisms Factors Affecting Transport
Oxygen Hemoglobin Dissolved in plasma pH, Temperature, PCO2, 2,3-BPG
Carbon Dioxide Bicarbonate ions (HCO3-) Carbaminohemoglobin, Dissolved CO2 PO2, pH, Carbonic anhydrase activity

Conclusion

Gaseous transport in blood is a complex and highly regulated process essential for life. The efficient delivery of oxygen and removal of carbon dioxide rely on the unique properties of hemoglobin, the bicarbonate buffering system, and the coordinated function of the respiratory and circulatory systems. Disruptions in any of these components can lead to significant physiological consequences. Further research continues to refine our understanding of these processes and develop strategies to address related clinical challenges.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Partial Pressure
The pressure exerted by a single gas in a mixture of gases. It is measured in millimeters of mercury (mmHg) or kilopascals (kPa).
Bohr Effect
The Bohr effect describes the shift in the oxygen dissociation curve caused by changes in pH or PCO2. Lower pH and higher PCO2 decrease hemoglobin's affinity for oxygen.

Key Statistics

Approximately 1.5% of oxygen is transported dissolved in plasma.

Source: Guyton and Hall Textbook of Medical Physiology (Knowledge cutoff 2023)

Approximately 70% of carbon dioxide is transported in the blood as bicarbonate ions.

Source: Ganong's Review of Medical Physiology (Knowledge cutoff 2023)

Examples

High Altitude Adaptation

Individuals acclimatizing to high altitudes experience increased erythropoiesis, leading to higher hemoglobin concentrations and improved oxygen-carrying capacity. They also exhibit increased 2,3-BPG levels, facilitating oxygen release to tissues.

Frequently Asked Questions

What is the Haldane effect?

The Haldane effect describes the phenomenon where decreased oxygen levels in blood promote the release of carbon dioxide from hemoglobin, enhancing CO2 transport from tissues to the lungs.

Topics Covered

BiologyPhysiologyRespirationBlood CirculationHemoglobin