UPSC MainsZOOLOGY-PAPER-II201710 Marks
Q3.

Explain in brief the role of 'isolation' in evolution by giving suitable examples.

How to Approach

This question requires a detailed explanation of the evolutionary significance of isolation. The answer should define isolation (both geographic and reproductive), explain how it leads to allopatric and sympatric speciation, and provide concrete examples. Structure the answer by first defining isolation, then detailing geographic isolation and its effects, followed by reproductive isolation and its mechanisms, and finally, concluding with the overall importance of isolation in driving biodiversity. Focus on clarity and use specific examples to illustrate the concepts.

Model Answer

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Introduction

Evolution, at its core, is driven by changes in the genetic makeup of populations over time. A crucial factor facilitating these changes is ‘isolation’ – the separation of populations, preventing or hindering gene flow. This separation can occur through various mechanisms, leading to divergent evolution and ultimately, the formation of new species. The concept of isolation was central to Darwin’s theory of evolution and continues to be a cornerstone of modern evolutionary biology. Understanding the role of isolation is fundamental to comprehending the incredible biodiversity observed on Earth.

Geographic Isolation (Allopatric Speciation)

Geographic isolation, also known as allopatric speciation, occurs when populations are physically separated by a geographic barrier, such as mountains, rivers, oceans, or deserts. This prevents interbreeding and allows each population to evolve independently under different selective pressures.

  • Mechanism: The barrier restricts gene flow, leading to genetic divergence through mutation, natural selection, and genetic drift.
  • Example 1: Darwin’s Finches: The finches on the Galapagos Islands are a classic example. A single ancestral finch species arrived from the mainland, and subsequent geographic isolation on different islands led to the evolution of 13 distinct species, each adapted to a specific food source.
  • Example 2: Snapping Shrimp in Panama: The Isthmus of Panama rose approximately 3 million years ago, dividing populations of snapping shrimp in the Atlantic and Pacific Oceans. These populations have since diverged genetically and behaviorally, becoming distinct species.

Reproductive Isolation (Sympatric Speciation)

Reproductive isolation occurs when populations live in the same geographic area (sympatric) but cannot interbreed due to various biological factors. This can lead to sympatric speciation – the evolution of new species from within the same geographic region.

Prezygotic Barriers

These barriers prevent mating or fertilization from occurring.

  • Habitat Isolation: Populations live in different habitats within the same area.
  • Temporal Isolation: Populations breed during different times of day or year.
  • Behavioral Isolation: Differences in courtship rituals or other behaviors prevent mate recognition.
  • Mechanical Isolation: Physical differences prevent successful mating.
  • Gametic Isolation: Eggs and sperm are incompatible.

Postzygotic Barriers

These barriers occur after the formation of a hybrid zygote.

  • Reduced Hybrid Viability: Hybrid offspring are unable to develop or survive.
  • Reduced Hybrid Fertility: Hybrid offspring are sterile.
  • Hybrid Breakdown: First-generation hybrids are fertile, but subsequent generations are infertile.

Example 1: Apple Maggot Flies: Originally, these flies laid their eggs only on hawthorn fruits. However, after the introduction of apples to North America, some flies began to lay their eggs on apples. This led to reproductive isolation as flies specializing on hawthorns continued to mate on hawthorn trees, while those specializing on apples mated on apple trees, eventually leading to genetic divergence.

Example 2: Cichlid Fishes in African Lakes: The diverse cichlid fish species in African lakes like Victoria and Malawi have undergone rapid sympatric speciation, driven by sexual selection and specialization on different food sources. Differences in coloration and mating calls contribute to reproductive isolation.

The Role of Genetic Drift and Natural Selection

Isolation, whether geographic or reproductive, sets the stage for genetic drift and natural selection to operate independently on isolated populations. Genetic drift, particularly strong in small populations, can lead to random changes in allele frequencies. Natural selection, acting on different environmental conditions, favors different traits in each population, further accelerating divergence. Over time, these accumulated genetic differences can become so significant that the populations are no longer able to interbreed, even if the isolating barrier is removed, resulting in the formation of distinct species.

Type of Isolation Mechanism Example
Geographic (Allopatric) Physical barrier prevents gene flow Darwin’s Finches
Reproductive (Sympatric) Biological factors prevent interbreeding Apple Maggot Flies

Conclusion

Isolation, in its various forms, is a fundamental driver of evolution and biodiversity. By restricting gene flow, it allows populations to diverge genetically and adapt to different environments, ultimately leading to the formation of new species. Understanding the mechanisms of isolation is crucial for conservation efforts, as fragmented habitats and reduced population sizes can exacerbate the effects of isolation and increase the risk of extinction. Continued research into the complexities of isolation will further refine our understanding of the evolutionary processes shaping life on Earth.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Speciation
The evolutionary process by which new biological species arise.
Gene Flow
The transfer of genetic variation from one population to another.

Key Statistics

Approximately 98% of all species that have ever lived are now extinct (based on fossil records as of 2023).

Source: Raup, D. M. (1991). Extinction: Bad luck and bad genes.

It is estimated that there are between 8.7 million and 1 trillion species on Earth (based on 2011 estimates).

Source: Mora, C., et al. (2011). How many species are there on Earth and in the ocean?

Examples

The Isthmus of Panama

The formation of the Isthmus of Panama approximately 3 million years ago dramatically altered marine ecosystems, leading to allopatric speciation in numerous marine organisms, including snapping shrimp and fish.

Frequently Asked Questions

Can isolation occur without a physical barrier?

Yes, reproductive isolation can occur even when populations live in the same geographic area. This is known as sympatric speciation and is driven by biological factors that prevent interbreeding.

Topics Covered

BiologyEvolutionSpeciationGenetic DriftNatural Selection