UPSC MainsBOTANY-PAPER-I201810 Marks
Q22.

Describe the different types of stomata in dicots.

How to Approach

This question requires a detailed understanding of plant anatomy, specifically focusing on stomatal types found in dicotyledonous plants. The answer should systematically describe each type – anomocytic, anisocytic, paracytic, and diacytic – outlining their structural characteristics, distribution, and any functional implications. A comparative approach, potentially using a table, would be beneficial. Diagrams, while not directly requested, are implicitly expected in a botany answer and should be mentally visualized while describing the structures.

Model Answer

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Introduction

Stomata are microscopic pores present on the epidermis of leaves and stems, crucial for gaseous exchange – facilitating carbon dioxide uptake for photosynthesis and oxygen release. Their structure and regulation are vital for plant survival and adaptation. While stomata are ubiquitous, their morphology varies significantly across plant species. In dicotyledonous plants, four primary types of stomata are recognized, classified based on the arrangement of surrounding subsidiary cells. Understanding these variations is fundamental to comprehending plant physiology and ecological adaptations. This answer will detail the characteristics of each stomatal type found in dicots: anomocytic, anisocytic, paracytic, and diacytic.

Anomocytic Stomata

Also known as irregular-celled stomata, anomocytic stomata are the most primitive and common type. They are characterized by the guard cells being surrounded by a variable number of epidermal cells, which are morphologically indistinguishable from other epidermal cells. There is no specific arrangement or pattern to the surrounding cells; they appear randomly distributed.

  • Structure: Guard cells are kidney-shaped and surrounded by irregularly shaped epidermal cells.
  • Distribution: Found in many dicot families, including Solanaceae (e.g., tomato, potato) and Cucurbitaceae (e.g., cucumber, watermelon).
  • Function: Generally considered less efficient in regulating transpiration due to the lack of specialized subsidiary cells.

Anisocytic Stomata

Anisocytic stomata, or unequal-celled stomata, are distinguished by the presence of three subsidiary cells surrounding the guard cells. Crucially, one subsidiary cell is significantly larger than the other two. This asymmetry is the defining feature of this type.

  • Structure: Guard cells are flanked by three subsidiary cells, with one being markedly larger and more elongated than the others.
  • Distribution: Commonly found in the families Cruciferae (Brassicaceae – e.g., mustard, cabbage) and Boraginaceae (e.g., borage, forget-me-not).
  • Function: The differing sizes of the subsidiary cells may contribute to more efficient stomatal closure and opening, potentially offering better control over gas exchange.

Paracytic Stomata

Paracytic stomata, also called parallel-celled stomata, are characterized by the presence of two parallel subsidiary cells flanking the guard cells. These subsidiary cells are aligned parallel to the longitudinal axis of the guard cells.

  • Structure: Guard cells are bordered by two parallel subsidiary cells, which are typically rectangular in shape.
  • Distribution: Predominantly found in the family Fabaceae (Leguminosae – e.g., pea, bean) and Oleaceae (e.g., olive, jasmine).
  • Function: The parallel arrangement of subsidiary cells is thought to facilitate efficient regulation of stomatal aperture, potentially enhancing water use efficiency.

Diacytic Stomata

Diacytic stomata, or dumbbell-shaped stomata, are characterized by the presence of two subsidiary cells that are perpendicular to the longitudinal axis of the guard cells. These cells appear to encircle the guard cells, giving the stomata a dumbbell-like appearance.

  • Structure: Guard cells are surrounded by two subsidiary cells positioned at right angles to the guard cell’s long axis.
  • Distribution: Commonly found in the families Euphorbiaceae (e.g., castor bean, poinsettia) and Asteraceae (Compositae – e.g., sunflower, daisy).
  • Function: The unique arrangement of subsidiary cells may provide enhanced structural support and contribute to precise control of stomatal movement.

The following table summarizes the key differences between the four stomatal types:

Stomatal Type Subsidiary Cell Arrangement Distribution (Example Families)
Anomocytic Irregular, variable number Solanaceae, Cucurbitaceae
Anisocytic Three, one significantly larger Cruciferae, Boraginaceae
Paracytic Two, parallel Fabaceae, Oleaceae
Diacytic Two, perpendicular Euphorbiaceae, Asteraceae

Conclusion

In conclusion, the four types of stomata – anomocytic, anisocytic, paracytic, and diacytic – represent variations in stomatal structure adapted to different environmental conditions and plant families. These differences in subsidiary cell arrangement influence stomatal regulation and, consequently, plant water use efficiency and gas exchange. Understanding these variations is crucial for comprehending plant adaptation and responses to changing climates. Further research into the functional significance of each type could contribute to developing strategies for improving crop resilience and productivity.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Subsidiary Cells
Epidermal cells surrounding the guard cells of a stoma, playing a role in regulating stomatal aperture.
Transpiration
The process by which water is carried through plants and evaporated from aerial parts, such as leaves, stems and flowers. Stomata play a key role in regulating transpiration rates.

Key Statistics

Approximately 90% of water loss in plants occurs through stomata.

Source: Raven, P. H., Evert, R. F., & Eichhorn, S. E. (2013). Biology of Plants (8th ed.). W. H. Freeman and Company.

Stomatal conductance, a measure of the rate of carbon dioxide entering or water vapor exiting through stomata, can vary by a factor of 100 depending on environmental conditions.

Source: Jones, H. G. (1992). Plants and Microclimate. Cambridge University Press.

Examples

Sunflower (Helianthus annuus)

Sunflowers exhibit diacytic stomata, which are believed to contribute to their ability to thrive in relatively dry conditions by regulating water loss.

Frequently Asked Questions

Do all plants have the same type of stomata?

No, stomatal types vary significantly between plant species and even within different parts of the same plant. The type often correlates with the plant's habitat and evolutionary history.

Topics Covered

BotanyPlant AnatomyPhysiologyStomatal StructureTranspirationGas Exchange