UPSC MainsAGRICULTURE-PAPER-I201910 Marks150 Words
Q16.

What is polyploidy? Describe its application in crop improvement with its limitations.

How to Approach

This question requires a clear understanding of polyploidy and its application in crop improvement. The approach should begin by defining polyploidy and explaining its different types. The body should then detail the benefits of polyploidy in crop improvement, followed by a discussion of its limitations. A structured approach with clear headings and subheadings will ensure a comprehensive and well-organized answer. Specific examples of crops improved through polyploidy will strengthen the response.

Model Answer

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Introduction

Polyploidy, a fascinating phenomenon in genetics, refers to the condition where an organism possesses more than two complete sets of chromosomes. Unlike normal diploid organisms (2n) having two sets, polyploids can have 3n (triploid), 4n (tetraploid), or even higher numbers. This condition arises due to errors during cell division, specifically meiosis. While initially considered a genetic defect, polyploidy has been harnessed as a powerful tool in crop improvement, leading to significant advancements in yield, size, and disease resistance. The discovery of induced mutations and plant breeding techniques has further enhanced the application of polyploidy in agriculture.

What is Polyploidy?

Polyploidy occurs when there's a duplication of the entire chromosome set. It's broadly classified into:

  • Autopolyploidy: Duplication of chromosomes from a single species (e.g., 4n from 2n).
  • Allopolyploidy: Combination of chromosome sets from two different species (e.g., 6n from 3n + 3n).

This differs significantly from aneuploidy, which involves a change in the number of individual chromosomes, not entire sets.

Application of Polyploidy in Crop Improvement

Polyploidy offers several advantages for crop improvement:

  • Increased Size & Yield: Polyploid plants generally exhibit larger cell size, leading to increased fruit, seed, or overall plant size. This directly translates to higher yields.
  • Enhanced Genetic Diversity: Allopolyploidy introduces new genetic combinations, creating greater genetic diversity within a species. This is valuable for breeding programs.
  • Improved Disease Resistance: Polyploidy can sometimes confer resistance to certain diseases due to the presence of multiple copies of disease resistance genes.
  • Increased Vigor (Heterosis): In allopolyploids, the combination of genes from different species can lead to hybrid vigor (heterosis).

Examples:

  • Wheat: Modern bread wheat (Triticum aestivum) is an allopolyploid (AABBDD), derived from three different species.
  • Cotton: Many cultivated cotton varieties are tetraploid.
  • Potato: Most commercial potato varieties are tetraploid, exhibiting larger tubers compared to their diploid counterparts.
  • Banana: The commercially grown banana (Cavendish) is triploid, resulting in seedless fruits.

Limitations of Polyploidy in Crop Improvement

Despite its advantages, polyploidy has limitations:

  • Sterility: Odd-numbered polyploids (e.g., triploids – 3n) are often sterile due to uneven chromosome pairing during meiosis. This is because chromosomes struggle to find proper partners, leading to non-viable gametes.
  • Complex Genome: Polyploid genomes are complex, making genetic analysis and breeding more challenging. Identifying and manipulating specific genes is difficult.
  • Reduced Fertility in Subsequent Generations: While initial generations might show increased vigor, fertility can decline in subsequent generations due to genetic instability.
  • Morphological Instability: Polyploids can sometimes exhibit unpredictable morphological changes, making it difficult to maintain desired traits.
  • Increased Maintenance Costs: Larger plant size, a common outcome of polyploidy, can increase maintenance costs like irrigation and fertilization.

The use of colchicine, a chemical that inhibits spindle formation and induces chromosome doubling, is a common method for creating polyploids. However, this process can be laborious and may result in undesirable mutations.

Type Chromosome Number Fertility Example
Diploid (2n) 2 sets Fertile Most naturally occurring plants
Triploid (3n) 3 sets Often Sterile Banana (Cavendish)
Tetraploid (4n) 4 sets Generally Fertile Potato

Conclusion

Polyploidy represents a significant evolutionary force and a valuable tool in crop improvement. While offering benefits like increased size, yield, and genetic diversity, its limitations, particularly sterility in odd-numbered polyploids and genome complexity, require careful consideration. Future research focusing on overcoming these limitations, such as through genome editing techniques, could further enhance the application of polyploidy in developing improved crop varieties to address global food security challenges.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Aneuploidy
A condition where the number of individual chromosomes is altered (e.g., having one extra or one missing chromosome).
Colchicine
A chemical compound used to induce polyploidy by disrupting spindle formation during cell division.

Key Statistics

Approximately 30-50% of plant species are polyploid.

Source: Knowledge cutoff - widely accepted estimate

Triploid plants, due to their sterility, often require vegetative propagation for commercial cultivation.

Source: Knowledge cutoff - general observation

Examples

Seedless Watermelon

Triploid watermelons are produced to create seedless varieties. They are sterile and require pollination from a diploid watermelon.

Frequently Asked Questions

Can polyploidy occur naturally?

Yes, polyploidy can arise naturally through errors in meiosis during gamete formation or through chromosome duplication events.

Topics Covered

AgricultureGeneticsPlant BreedingPolyploidyCrop ImprovementPlant Genetics