UPSC MainsAGRICULTURE-PAPER-II201920 Marks
Q9.

Explain the role of plant growth substances in the various developmental stages of a plant with special reference to horticultural crops.

How to Approach

This question requires a comprehensive understanding of plant growth substances (phytohormones) and their roles in plant development. A structured approach is crucial, beginning with a definition and classification of phytohormones. The answer should then detail the roles of each hormone in different developmental stages (seed germination, vegetative growth, flowering, fruit development, senescence), with specific examples relevant to horticultural crops. Finally, it should briefly touch upon the practical applications of phytohormones in horticulture. The structure should follow a logical flow, with clear headings and subheadings.

Model Answer

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Introduction

Plant growth substances, also known as phytohormones, are naturally occurring organic compounds that regulate various aspects of plant growth and development. These signaling molecules, present in minute quantities, orchestrate processes ranging from seed germination to senescence. The discovery of auxin in 1930s revolutionized our understanding of plant physiology. The increasing demand for high-quality horticultural crops necessitates a thorough understanding of these hormones, enabling targeted interventions to enhance yield and quality. This response will detail the role of key phytohormones in different developmental stages, focusing on relevance to horticultural practices.

Plant Growth Substances: An Overview

Phytohormones don't act in isolation; their effects are often synergistic or antagonistic, influencing each other's actions. They are broadly classified into several categories:

  • Auxins: Primarily involved in cell elongation, apical dominance, and root initiation.
  • Gibberellins: Promote stem elongation, seed germination, and flowering.
  • Cytokinins: Stimulate cell division, delay senescence, and influence apical dominance.
  • Abscisic Acid (ABA): Involved in stress responses (drought, salinity), seed dormancy, and abscission.
  • Ethylene: Promotes fruit ripening, senescence, and abscission.
  • Brassinosteroids: Influence cell elongation, vascular differentiation, and stress tolerance.
  • Jasmonates: Involved in defense responses, senescence, and fruit ripening.
  • Salicylic Acid: Plays a role in systemic acquired resistance (SAR) against pathogens.

Role of Phytohormones in Different Developmental Stages

1. Seed Germination

Gibberellins play a critical role in overcoming seed dormancy and promoting germination. They stimulate the synthesis of hydrolytic enzymes that break down stored food reserves within the seed. Abscisic Acid (ABA), conversely, maintains seed dormancy under unfavorable conditions.

Example: Gibberellic acid treatment is often used to break dormancy in fruit tree seeds like apples and peaches, improving germination rates.

2. Vegetative Growth

Auxins primarily promote cell elongation and apical dominance, influencing shoot growth. Cytokinins stimulate cell division and counter apical dominance, promoting lateral bud growth. The auxin:cytokinin ratio is a key determinant of branching patterns.

Statistics: Studies show that increasing cytokinin levels in young seedlings can lead to a bushier growth habit, increasing the leaf area index (LAI) which is crucial for photosynthesis in horticultural crops.

3. Flowering

Gibberellins induce flowering in long-day plants under non-inductive conditions. Auxins also play a role in flower initiation and development. The timing of flowering is crucial in horticultural crops to ensure optimal harvest periods.

Scheme: The Pradhan Mantri Kisan SAMPADA Yojana (PMKSY) includes components aimed at improving horticulture production, which can indirectly benefit from optimized flowering through proper management of plant growth regulators.

4. Fruit Development and Ripening

Auxins are produced in developing fruits and contribute to cell enlargement and fruit set. Ethylene is the primary hormone responsible for fruit ripening, triggering changes in color, texture, and flavor. Jasmonates also contribute to ripening and flavor development.

Case Study: The Cavendish banana, the most widely cultivated banana variety globally, is highly susceptible to Panama disease. Research is focusing on understanding the role of jasmonates in banana defense mechanisms against fungal pathogens to develop disease-resistant varieties.

5. Senescence and Abscission

Ethylene promotes senescence (aging) in leaves and flowers, leading to chlorophyll degradation and yellowing. Abscisic Acid (ABA) is involved in abscission (leaf and fruit drop) by triggering the formation of an abscission layer.

Definition: Senescence is the process of aging in plants, characterized by a decline in metabolic activity and eventual death.

Practical Applications in Horticultural Crops

Horticultural practices often utilize phytohormones to manipulate plant development:

  • Rooting Cuttings: Auxins (e.g., IBA, NAA) are used to stimulate root formation in stem cuttings for vegetative propagation.
  • Fruit Thinning: Auxins are employed to thin out fruit clusters in apple and pear orchards, improving fruit size and quality.
  • Delaying Ripening: Ethylene inhibitors are used to extend the shelf life of fruits like tomatoes and bananas.
  • Promoting Flowering: Gibberellins are used to induce flowering in pineapple plants.
Hormone Primary Role Horticultural Application
Auxin Cell elongation, root initiation Rooting cuttings, fruit thinning
Gibberellin Stem elongation, seed germination Inducing flowering in pineapples
Cytokinin Cell division, delay senescence Promoting bushier growth
Ethylene Fruit ripening, senescence Extending shelf life of fruits

FAQ

FAQ: Can phytohormones be harmful if used excessively?

Answer: Yes, excessive use of phytohormones can lead to undesirable effects like abnormal growth, fruit drop, and physiological disorders. Careful calibration and application based on crop needs are essential.

Conclusion

Plant growth substances are critical regulators of plant development, impacting various stages from germination to senescence. Understanding their individual roles and interactions is crucial for optimizing horticultural practices and enhancing crop yield and quality. Continued research into the complexities of phytohormone signaling pathways holds the potential to further refine horticultural techniques and develop more resilient and productive crop varieties. The judicious use of these regulators, coupled with sustainable agricultural practices, is vital for ensuring food security and environmental sustainability.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Apical Dominance
The phenomenon where the main shoot of a plant grows faster than other side shoots, suppressing their growth.
Abscission
The natural shedding of organs (leaves, flowers, fruits) from a plant.

Key Statistics

The global horticulture market is estimated to be worth over $300 billion, highlighting the economic importance of optimizing plant growth and development (Source: FAO, 2023 - Knowledge cutoff).

Source: FAO

Ethylene production in ripening tomatoes increases dramatically, reaching levels up to 100 ppm (parts per million) (Source: Knowledge cutoff).

Source: Plant Physiology textbook

Examples

Pineapple Flowering

Pineapple plants typically flower after several years. Application of gibberellic acid can induce flowering in pineapple plants much earlier, significantly reducing the time to harvest.

Frequently Asked Questions

What is the difference between natural and synthetic phytohormones?

Natural phytohormones are produced by the plant itself. Synthetic phytohormones are chemically synthesized to mimic the effects of natural hormones. Both can be used in horticulture, but synthetic forms often have higher potency.

Topics Covered

BotanyHorticulturePlant PhysiologyPlant HormonesPlant DevelopmentHorticultural Crops