UPSC MainsBOTANY-PAPER-I202215 Marks150 Words
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Q14.

Give a detailed account of types of stomata in angiosperms.

How to Approach

This question requires a detailed understanding of stomatal types found in angiosperms. The answer should focus on describing the different types – cydonia, anomocytic, dicotyledonous, and actinocytic – along with their characteristic features and occurrence. A comparative approach, potentially using a table, would be beneficial. Mentioning the functional significance of each type will add value. The answer should be concise, sticking to the 150-word limit.

Model Answer

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Introduction

Stomata are microscopic pores present on the epidermis of leaves and stems, crucial for gaseous exchange and transpiration in plants. Their structure varies considerably across different plant species, particularly within angiosperms. These variations are categorized into distinct types based on the surrounding subsidiary cells. Understanding these stomatal types provides insights into plant adaptation and evolutionary relationships. The classification, initially proposed by Metcalfe and Chalkley, helps in plant taxonomic studies and understanding physiological processes.

Types of Stomata in Angiosperms

Angiosperms exhibit four main types of stomata:

1. Cydonia Type (Apple Type)

  • Surrounded by two subsidiary cells flanking the guard cells.
  • Subsidiary cells are kidney-shaped.
  • Common in members of the family Rosaceae (e.g., Apple, Pear).

2. Anomocytic Type (Random Type)

  • Guard cells are surrounded by a variable number of subsidiary cells, appearing randomly arranged.
  • Most primitive type.
  • Found in dicotyledonous plants like Solanum (Potato) and Arabidopsis.

3. Dicotyledonous Type (Rubiaceous Type)

  • Guard cells are surrounded by two distinct, regularly arranged subsidiary cells.
  • Subsidiary cells are rectangular.
  • Common in families like Rubiaceae (e.g., Coffee, Cinchona).

4. Actinocytic Type (Cruciferous Type)

  • Guard cells are surrounded by several radiating subsidiary cells.
  • Subsidiary cells appear star-shaped.
  • Characteristic of the family Brassicaceae (e.g., Mustard, Radish).
Stomatal Type Subsidiary Cell Arrangement Occurrence
Cydonia Two kidney-shaped cells Rosaceae
Anomocytic Variable, random Solanum, Arabidopsis
Dicotyledonous Two rectangular cells Rubiaceae
Actinocytic Radiating, star-shaped Brassicaceae

Conclusion

In conclusion, the diversity in stomatal types within angiosperms reflects adaptations to varying environmental conditions and evolutionary histories. Each type, characterized by its unique arrangement of subsidiary cells, plays a vital role in regulating gas exchange and water loss. Studying these variations is crucial for understanding plant physiology, taxonomy, and responses to environmental stress. Further research into the genetic control of stomatal development could lead to strategies for improving crop resilience.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Subsidiary Cells
Epidermal cells surrounding the guard cells of a stoma, contributing to its structure and function.
Stomatal Conductance
A measure of the rate of carbon dioxide entering or water vapor exiting through the stomata, reflecting the degree of stomatal opening.

Key Statistics

Approximately 60% of terrestrial photosynthesis is regulated by stomatal conductance (source: Leuning, 1995, Plant, Cell & Environment).

Source: Leuning, R. (1995). A critical appraisal of a model for stomatal conductance. *Plant, Cell & Environment*, *18*(2), 241–252.

Stomatal closure can reduce transpiration rates by up to 90% in some plant species during periods of water stress (source: Kramer, 1983, Water Relations of Plants).

Source: Kramer, P. J. (1983). *Water relations of plants*. Academic Press.

Examples

Drought Tolerance in <em>Nerium oleander</em>

<em>Nerium oleander</em> (Oleander) exhibits sunken stomata and a thick cuticle, reducing water loss in arid environments. This is an example of structural adaptation related to stomatal function.

Frequently Asked Questions

What is the functional significance of different stomatal types?

Different stomatal types may exhibit varying degrees of sensitivity to environmental cues like light, CO2 concentration, and water stress, influencing their opening and closing mechanisms and thus, the plant's physiological response.

Topics Covered

BotanyPlant AnatomyPlant PhysiologyTranspirationGas Exchange