UPSC MainsZOOLOGY-PAPER-I202215 Marks
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Q14.

Describe habitat, habits and general features of Herdmania. Write a note on its development.

How to Approach

This question requires a detailed understanding of *Herdmania*, a urochordate. The answer should be structured into three main parts: habitat and habits, general features (morphology, physiology, and anatomy), and its development. Focus on key characteristics that distinguish it from other chordates. Include diagrams where appropriate (though not possible in text-only format, mention their relevance). The development section should cover fertilization, cleavage, gastrulation, and metamorphosis. A comparative approach highlighting its unique features will enhance the answer.

Model Answer

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Introduction

*Herdmania*, commonly known as the sea squirt, is a sessile, marine urochordate belonging to the subphylum Tunicata. These filter-feeding organisms are found attached to rocks, shells, or other submerged surfaces in shallow coastal waters worldwide. Their simple body plan and unique developmental stages make them a fascinating subject for zoological study, offering insights into the evolution of chordates. Understanding *Herdmania*’s biology is crucial for comprehending the broader context of invertebrate zoology and chordate phylogeny. This answer will detail its habitat, habits, general features, and developmental stages.

Habitat and Habits

Herdmania typically inhabits shallow, clear, and relatively calm marine environments. They are commonly found attached to rocks, shells, pilings, and other hard substrates in intertidal and subtidal zones. Their distribution is influenced by factors like salinity, temperature, and water currents. They are cosmopolitan in distribution, found in temperate and tropical seas.

  • Attachment: *Herdmania* attaches to the substrate via a stalk secreted by the larval stage.
  • Feeding: They are filter feeders, drawing water in through the incurrent siphon, filtering out phytoplankton and organic particles, and expelling water through the excurrent siphon.
  • Sessile Lifestyle: Adult *Herdmania* are sessile, remaining permanently attached to their substrate.
  • Solitary or Colonial: They can exist as solitary individuals or form small colonies.

General Features

Morphology

The adult *Herdmania* exhibits a simple, sac-like body plan. It is covered by a tough, gelatinous tunic, hence the name Tunicata. Key features include:

  • Tunic: A cellulose-rich outer covering providing protection.
  • Siphons: Two openings – the incurrent siphon (for water intake) and the excurrent siphon (for waste expulsion).
  • Branchial Basket: An internal structure containing gill slits used for respiration and filter feeding.
  • Atrium: The cavity surrounding the branchial basket.
  • Visceral Mass: Contains the digestive, reproductive, and excretory organs.

Physiology

*Herdmania* exhibits several unique physiological adaptations:

  • Circulatory System: An open circulatory system with a dorsal heart that pumps hemolymph (blood) through sinuses.
  • Respiratory System: Gas exchange occurs through the gill slits in the branchial basket.
  • Nervous System: A simple nervous system consisting of a cerebral ganglion and a visceral nerve.
  • Excretory System: Excretion occurs through the nephridia, which filter waste products from the blood.

Anatomy

The internal anatomy of *Herdmania* is relatively simple. The body cavity is spacious and contains the visceral organs. The notochord, a defining feature of chordates, is present only in the larval stage and is lost during metamorphosis. The presence of a post-anal tail is also limited to the larval stage.

Feature Description
Notochord Present in larva, absent in adult
Post-anal Tail Present in larva, absent in adult
Gill Slits Numerous, used for filter feeding and respiration
Tunic Protective outer covering made of cellulose

Development

The development of *Herdmania* is characterized by a unique life cycle involving metamorphosis. It includes the following stages:

  • Fertilization: External fertilization occurs in the water column.
  • Cleavage: Radial and determinate cleavage leads to the formation of a blastula.
  • Gastrulation: Gastrulation results in the formation of a gastrula with an archenteron (primitive gut).
  • Larval Stage: A free-swimming larva emerges, possessing a notochord, post-anal tail, and a well-developed nervous system – exhibiting chordate characteristics.
  • Metamorphosis: The larva settles on a substrate, undergoes metamorphosis, and transforms into the sessile adult form. During metamorphosis, the tail and notochord are resorbed, and the siphons and branchial basket develop.

Conclusion

In conclusion, *Herdmania* represents a fascinating example of a urochordate, showcasing a unique blend of invertebrate and chordate characteristics. Its simple body plan, filter-feeding habits, and distinctive developmental stages provide valuable insights into the evolution of chordates. The loss of the notochord and post-anal tail during metamorphosis highlights the dramatic changes that occur during its life cycle. Further research into *Herdmania* and other urochordates will continue to refine our understanding of chordate phylogeny and development.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Urochordata
The subphylum Urochordata, also known as tunicates, are marine invertebrates characterized by a notochord present only in the larval stage and a tunic covering the body.
Metamorphosis
A biological process by which an animal physically develops after birth or hatching, involving a conspicuous and relatively sudden change in the animal's body structure through cell growth and differentiation.

Key Statistics

Approximately 3,000 species of tunicates, including *Herdmania*, have been identified worldwide.

Source: Barnes, R. S. K., Calow, P., Olive, P. J. W., Golding, D. W., & Spicer, J. I. (2001). The invertebrates: a synthesis. Blackwell Science.

Tunicates contribute significantly to benthic biomass in many coastal ecosystems, estimated to be around 5-10% in some regions.

Source: Knowledge cutoff - based on general zoological literature as of 2023.

Examples

Sea Squirt Farming

In some parts of Asia, particularly Korea and Japan, sea squirts (including species similar to *Herdmania*) are commercially farmed for human consumption. They are considered a delicacy.

Frequently Asked Questions

What is the significance of the larval stage in *Herdmania*?

The larval stage is significant because it exhibits all the defining characteristics of chordates – notochord, dorsal hollow nerve cord, pharyngeal slits, and post-anal tail. This stage provides evidence for the evolutionary relationship between urochordates and other chordates.

Topics Covered

ZoologyBiologyInvertebratesUrochordataHerdmania Anatomy