UPSC MainsZOOLOGY-PAPER-II202210 Marks150 Words
Q19.

Illustrate the counter-current mechanism by vasa recta and mention its physiological importance.

How to Approach

This question requires a detailed understanding of renal physiology, specifically the function of vasa recta in maintaining the concentration gradient within the kidney. The answer should begin by defining the counter-current mechanism and then explain how vasa recta contributes to it. Focus on the anatomical arrangement and its role in preventing washout of solutes from the medulla. Mention the physiological importance in terms of urine concentration and water reabsorption. A diagram, though not explicitly asked for, would enhance the answer.

Model Answer

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Introduction

The mammalian kidney exhibits a remarkable ability to produce urine that is significantly more concentrated than blood plasma. This is achieved through the counter-current mechanism, a complex interplay between the loop of Henle, vasa recta, and the collecting duct. The vasa recta, a specialized network of capillaries, plays a crucial role in maintaining the osmotic gradient established by the loop of Henle within the renal medulla. Understanding the counter-current exchange system facilitated by vasa recta is fundamental to comprehending renal function and overall fluid balance in the body.

The Counter-Current Mechanism & Vasa Recta

The counter-current mechanism relies on the opposing flow of fluids in adjacent tubules and blood vessels, creating and maintaining an osmotic gradient in the renal medulla. This gradient is essential for concentrating urine.

Anatomy of Vasa Recta

Vasa recta are long, hairpin-shaped capillaries that run parallel to the loops of Henle in the renal medulla. They originate from the efferent arterioles of the glomeruli and descend into the medulla, forming a network around the loops of Henle. They then ascend back towards the cortex. This unique anatomical arrangement is critical for their function.

Counter-Current Exchange in Vasa Recta

The vasa recta participate in a counter-current exchange system. As blood descends into the medulla, it encounters increasing solute concentrations. Due to the permeability of the vasa recta capillaries, water moves out of the blood and solutes move into the blood, minimizing the washout of solutes from the medulla. Conversely, as blood ascends, it encounters decreasing solute concentrations. Water moves into the blood, and solutes move out, maintaining the medullary gradient.

Key Features of Counter-Current Exchange:

  • Descending Limb: Blood gains solutes, loses water.
  • Ascending Limb: Blood loses solutes, gains water.
  • Low Blood Flow: The relatively low blood flow rate through the vasa recta is crucial. A high flow rate would disrupt the gradient by rapidly removing solutes.
  • Permeability: The capillaries of vasa recta are highly permeable to water and solutes.

Physiological Importance

The counter-current mechanism facilitated by vasa recta is vital for several physiological processes:

  • Urine Concentration: It allows the kidneys to produce concentrated urine, conserving water when the body is dehydrated.
  • Water Reabsorption: By maintaining the medullary gradient, it enhances water reabsorption from the collecting duct, further contributing to water conservation.
  • Regulation of Blood Osmolarity: It plays a key role in maintaining the body’s fluid and electrolyte balance.

Without the vasa recta’s counter-current exchange, the medullary gradient would be lost, and the kidneys would be unable to concentrate urine effectively, leading to excessive water loss and potential dehydration.

Component Role in Counter-Current Mechanism
Loop of Henle Establishes the initial osmotic gradient in the medulla.
Vasa Recta Maintains the medullary gradient by counter-current exchange.
Collecting Duct Reabsorbs water based on the medullary gradient, concentrating urine.

Conclusion

In conclusion, the vasa recta’s counter-current exchange system is an indispensable component of the mammalian kidney’s ability to regulate water balance and produce concentrated urine. Its unique anatomical arrangement and physiological function are crucial for maintaining the medullary osmotic gradient, ensuring efficient water reabsorption and preventing dehydration. Disruptions to this mechanism can lead to significant impairments in renal function and overall homeostasis.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Osmolarity
Osmolarity is a measure of solute concentration, specifically the number of osmoles of solute per liter of solvent (water). It reflects the ability of a solution to draw water across a semipermeable membrane.
Counter-current multiplier
The counter-current multiplier refers to the process by which the loop of Henle creates and maintains an osmotic gradient in the renal medulla, multiplying the effect of a small initial difference in solute concentration.

Key Statistics

The human kidney filters approximately 180 liters of fluid per day, but only about 1.5-2 liters are excreted as urine. The remaining 178-179 liters are reabsorbed, highlighting the kidney’s efficiency in water conservation (Source: Guyton and Hall Textbook of Medical Physiology, 13th Edition - Knowledge Cutoff 2023).

Source: Guyton and Hall Textbook of Medical Physiology

Approximately 10% of the global population is affected by chronic kidney disease (CKD) (Source: World Health Organization, 2021 - Knowledge Cutoff 2023).

Source: World Health Organization

Examples

Diabetes Insipidus

Diabetes insipidus is a condition characterized by the inability to concentrate urine due to a deficiency in antidiuretic hormone (ADH) or a resistance to its effects. This disrupts the counter-current mechanism and leads to excessive water loss, demonstrating the importance of this system in maintaining fluid balance.

Frequently Asked Questions

What would happen if the vasa recta were completely blocked?

If the vasa recta were completely blocked, the medullary gradient would rapidly dissipate as solutes would be washed out. This would severely impair the kidney’s ability to concentrate urine, leading to excessive water loss and potential dehydration. The kidney would function more like the kidney of a bird or fish, which lack a well-developed medullary gradient.

Topics Covered

BiologyPhysiologyKidney FunctionOsmoregulationRenal Physiology