UPSC MainsAGRICULTURE-PAPER-I202420 Marks
Q13.

What is the Arnon and Stout (1939) criteria for plant nutrient essentiality? Give account of forms of each essential plant nutrient element absorbed by plants.

How to Approach

This question requires a two-pronged approach. First, clearly explain the Arnon and Stout (1939) criteria for essentiality, highlighting the rationale behind each criterion. Second, systematically list each essential nutrient, detailing the forms in which plants absorb them – macronutrients (N, P, K, Ca, Mg, S) and micronutrients (Fe, Mn, Zn, Cu, B, Mo, Cl). Structure the answer using subheadings for clarity and employ tables where beneficial to present information concisely. Focus on factual accuracy and comprehensive coverage.

Model Answer

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Introduction

Plant nutrition is a cornerstone of agricultural productivity and ecosystem health. The availability of essential nutrients directly impacts plant growth, development, and ultimately, yield. Determining which elements are truly "essential" required a rigorous scientific framework. In 1939, Arnon and Stout established a definitive set of criteria to classify elements as essential for plant life. These criteria, still widely accepted today, provide a clear and objective basis for understanding nutrient requirements. This response will detail these criteria and outline the various forms in which plants absorb these vital elements.

Arnon and Stout (1939) Criteria for Plant Nutrient Essentiality

Arnon and Stout’s criteria provided a systematic approach to defining essentiality. These criteria are:

  • Essential for Growth: The element must be necessary for the plant to complete its life cycle (germination, growth, reproduction). Without the element, the plant cannot complete its life cycle.
  • Specific Role: The element must be involved in some specific metabolic or structural function within the plant. It cannot be substituted by another element.
  • Deficiency Symptoms: A deficiency in the element must produce characteristic morphological (observable) and/or physiological (functional) symptoms in the plant.
  • Reversal of Symptoms: The deficiency symptoms must be completely corrected by the addition of the element to the nutrient solution or soil.

These criteria ensure that an element’s essentiality is established through rigorous scientific observation and experimentation, eliminating elements that might be incidentally present but not fundamentally required for plant life.

Forms of Essential Plant Nutrients Absorbed by Plants

Essential plant nutrients are broadly classified into macronutrients (required in larger quantities) and micronutrients (required in smaller quantities). Plants absorb these nutrients primarily as ions, although the specific forms can vary depending on soil chemistry and plant species.

Macronutrients

Nutrient Role in Plant Absorption Forms
Nitrogen (N) Component of proteins, nucleic acids, chlorophyll; vital for vegetative growth NO3- (nitrate), NH4+ (ammonium)
Phosphorus (P) Energy transfer (ATP), nucleic acids, membrane structure H2PO4- (primary phosphate), HPO42- (secondary phosphate)
Potassium (K) Osmotic regulation, enzyme activation, stomatal control K+
Calcium (Ca) Cell wall structure, membrane stability, enzyme activation Ca2+
Magnesium (Mg) Chlorophyll molecule, enzyme activation Mg2+
Sulfur (S) Component of proteins, coenzymes SO42- (sulfate)

Micronutrients

Nutrient Role in Plant Absorption Forms
Iron (Fe) Chlorophyll synthesis, electron transport Fe2+, Fe3+ (depending on pH)
Manganese (Mn) Photosynthesis, enzyme activation Mn2+
Zinc (Zn) Enzyme activation, hormone regulation Zn2+
Copper (Cu) Enzyme activation, electron transport Cu2+
Boron (B) Cell wall synthesis, sugar transport B(OH)4- (borate)
Molybdenum (Mo) Nitrogen metabolism MoO42-
Chlorine (Cl) Photosynthesis, osmotic regulation Cl-

The availability of these nutrients in the soil is influenced by factors like pH, organic matter content, and microbial activity. Understanding these forms is crucial for effective fertilizer management and diagnosing nutrient deficiencies.

Case Study: Iron Deficiency Anemia in Rice Crops

In many regions, particularly those with alkaline soils, rice crops suffer from iron deficiency, manifesting as chlorosis (yellowing of leaves). This is because the availability of Fe2+ decreases in higher pH conditions. The application of chelated iron (Fe-EDTA) which are organic complexes of iron, improves iron uptake and corrects the deficiency. This highlights the importance of understanding nutrient forms and soil chemistry for effective management.

The Indian government's Soil Health Card Scheme (2015) aims to provide farmers with information about the nutrient status of their soils, facilitating targeted fertilizer application and promoting sustainable agriculture practices.

Conclusion

In conclusion, Arnon and Stout’s criteria provide a robust framework for identifying essential plant nutrients. These elements are absorbed in specific ionic forms, influenced by soil conditions and plant physiology. A thorough understanding of these criteria and absorption mechanisms is vital for optimizing plant nutrition, ensuring food security, and promoting sustainable agricultural practices. Further research focusing on improving nutrient use efficiency and reducing fertilizer losses remains crucial for environmental sustainability.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Chlorosis
Chlorosis is the process of losing chlorophyll from leaves or other photosynthetic parts of a plant, resulting in yellowing.
Chelation
Chelation is the process where metal ions are bound to organic molecules, creating soluble complexes that are more readily absorbed by plants.

Key Statistics

Approximately 30% of the world’s agricultural land is affected by nutrient deficiencies (FAO, knowledge cutoff).

Source: FAO

Approximately 60-80% of applied nitrogen is lost to the environment through volatilization, leaching, and denitrification (knowledge cutoff).

Source: Various agricultural research reports

Examples

Nitrogen Deficiency in Maize

Maize plants deficient in nitrogen exhibit stunted growth, pale green or yellow leaves, and reduced grain yield. This is a widespread issue in many agricultural regions.

Frequently Asked Questions

Why are micronutrients required in smaller amounts than macronutrients?

Micronutrients often function as cofactors for enzymes, meaning they are essential for enzyme activity but are needed in catalytic amounts rather than being structural components like macronutrients.