UPSC MainsBOTANY-PAPER-I202415 Marks150 Words
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Q25.

What are the causes of variability in regenerated plants in tissue cultures? Give an account of utility of such variants in improvement of crop plants with examples.

How to Approach

This question requires a detailed understanding of the mechanisms causing variability during plant tissue culture and how these variations can be exploited for crop improvement. The answer should begin by explaining the sources of variability, including genetic and epigenetic factors. Then, it should elaborate on the utility of these variants, providing specific examples of crops improved through tissue culture-induced variations. A structured approach, dividing the answer into causes of variability and its application, will be effective.

Model Answer

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Introduction

Plant tissue culture, a cornerstone of modern biotechnology, allows for the regeneration of whole plants from small pieces of plant tissue. However, regenerated plants often exhibit variability, differing from the parent plant. This variability arises due to a range of factors during the in vitro process. While often considered a limitation, this inherent variation provides a valuable resource for crop improvement, offering opportunities to select for desirable traits and accelerate breeding programs. Understanding the causes of this variability and harnessing its potential is crucial for enhancing agricultural productivity and adapting crops to changing environmental conditions.

Causes of Variability in Regenerated Plants

Variability in regenerated plants from tissue culture can be broadly categorized into genetic and epigenetic causes:

Genetic Causes

  • Somaclonal Variation: This refers to the genetic variation arising during the tissue culture process. It’s induced by mutations, chromosome rearrangements, and changes in ploidy levels. The high concentration of growth regulators, particularly auxins and cytokinins, can induce mutations.
  • Mutations: Exposure to mutagens (even inherent in the culture medium or due to irradiation) can lead to point mutations, deletions, or insertions in the plant genome.
  • Chromosome Aberrations: Tissue culture can induce chromosome breaks, translocations, and aneuploidy (abnormal number of chromosomes). These aberrations can lead to altered gene expression and phenotypic changes.
  • Genome Instability: Dedifferentiated cells in culture often exhibit genomic instability, making them prone to genetic changes.

Epigenetic Causes

  • DNA Methylation: Changes in DNA methylation patterns can alter gene expression without changing the underlying DNA sequence. These changes can be heritable.
  • Histone Modification: Alterations in histone proteins can affect chromatin structure and gene accessibility, leading to changes in gene expression.
  • Small RNA-mediated Gene Silencing: Small RNAs, such as microRNAs, can regulate gene expression by targeting mRNA for degradation or translational repression.

Utility of Variants in Crop Improvement

Somaclonal variation, despite its potential drawbacks, has proven to be a valuable tool for crop improvement. The following are some examples:

Examples of Crop Improvement through Tissue Culture Variants

  • Potato: Somaclonal variation has been extensively used in potato breeding to develop disease-resistant varieties. For example, several potato cultivars resistant to Potato Virus Y (PVY) were developed through tissue culture.
  • Rice: Tissue culture has been used to generate rice variants with improved yield, grain quality, and stress tolerance (e.g., salt tolerance).
  • Banana: Somaclonal variation has been utilized to develop banana cultivars with resistance to Fusarium wilt (Panama disease), a major threat to banana production.
  • Sugarcane: Tissue culture-derived sugarcane variants have been selected for higher sucrose content and improved disease resistance.
  • Tomato: Variants with altered fruit ripening characteristics and improved disease resistance have been obtained through tissue culture.

Strategies for Utilizing Somaclonal Variation

  • Selection of Superior Variants: Large populations of regenerated plants are screened for desirable traits.
  • Protoplast Fusion: Combining protoplasts from different varieties can create novel genetic combinations.
  • Genome Editing: Combining tissue culture with genome editing technologies (e.g., CRISPR-Cas9) can precisely introduce desired changes into the plant genome.

Table: Comparison of Genetic and Epigenetic Variations

Feature Genetic Variation Epigenetic Variation
Nature of Change Alteration in DNA sequence Change in gene expression without DNA sequence alteration
Stability Generally stable and heritable Can be unstable and reversible
Examples Mutations, chromosome rearrangements DNA methylation, histone modification

Conclusion

Variability in regenerated plants from tissue culture, stemming from both genetic and epigenetic sources, presents a significant opportunity for crop improvement. While somaclonal variation can be unpredictable, strategic screening and selection, coupled with advanced biotechnological tools like genome editing, can harness this variability to develop superior crop varieties with enhanced yield, disease resistance, and stress tolerance. Continued research into the mechanisms underlying somaclonal variation will further refine these techniques and accelerate crop breeding programs, contributing to global food security.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Somaclonal Variation
Genetic variation arising in plants regenerated from tissue culture. It is induced by mutations, chromosome rearrangements, and changes in ploidy levels during the in vitro process.
Epigenetics
The study of changes in gene expression that do not involve alterations to the underlying DNA sequence, but are heritable.

Key Statistics

Approximately 20% of new potato varieties released globally since 1980 have been derived from tissue culture-induced somaclonal variation.

Source: FAOSTAT, 2023 (based on knowledge cutoff)

The global tissue culture market is projected to reach USD 3.8 billion by 2028, growing at a CAGR of 9.8% from 2021.

Source: Market Research Future, 2021 (based on knowledge cutoff)

Examples

Development of Disease-Resistant Banana

The ‘Williams’ banana variety, highly susceptible to Fusarium wilt, was improved through tissue culture to generate variants resistant to the disease, saving the banana industry in several regions.

Frequently Asked Questions

Is somaclonal variation always beneficial?

No, somaclonal variation can also lead to undesirable traits. Therefore, careful screening and selection are crucial to identify and propagate only the superior variants.

Topics Covered

BotanyBiotechnologyTissue CulturePlant BreedingGenetic Variation