UPSC MainsZOOLOGY-PAPER-I202410 Marks150 Words
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Q2.

Metamorphosis in Insects

How to Approach

This question requires a detailed understanding of metamorphosis in insects. The answer should define metamorphosis, categorize its types (complete and incomplete), explain the hormonal control, and provide examples. A structured approach involving defining the process, detailing the types, outlining the hormonal regulation, and providing illustrative examples will be effective. Focus on biological mechanisms and evolutionary advantages.

Model Answer

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Introduction

Metamorphosis, derived from the Greek words ‘meta’ (change) and ‘morphe’ (form), refers to a biological process by which an animal physically develops after birth or hatching, involving a conspicuous and relatively sudden change in the animal's body structure through cell growth and differentiation. In insects, it is a remarkable adaptation that allows them to exploit different ecological niches during their life cycle, reducing intraspecific competition. This transformation is regulated by complex hormonal interactions and is crucial for insect survival and diversification. Understanding metamorphosis is fundamental to comprehending insect biology and its ecological role.

Types of Metamorphosis

Insect metamorphosis is broadly categorized into two main types:

1. Complete Metamorphosis (Holometabolism)

  • This involves four distinct life stages: egg, larva, pupa, and adult.
  • The larval stage is drastically different from the adult stage in morphology, feeding habits, and habitat.
  • A pupal stage, often enclosed in a cocoon or chrysalis, facilitates the dramatic reorganization of tissues.
  • Examples: Butterflies, moths, beetles, flies, bees, and wasps.

2. Incomplete Metamorphosis (Hemimetabolism)

  • This involves three life stages: egg, nymph, and adult.
  • The nymph resembles the adult but lacks fully developed wings and reproductive organs.
  • Nymphs undergo a series of molts (ecdysis), gradually developing into adults.
  • There is no distinct pupal stage.
  • Examples: Grasshoppers, dragonflies, cockroaches, termites, and true bugs.

Hormonal Control of Metamorphosis

Metamorphosis is primarily regulated by two key hormones:

1. Juvenile Hormone (JH)

  • Produced by the corpora allata.
  • High levels of JH during molting promote larval or nymphal development.
  • Decreasing levels of JH trigger pupation (in holometabolous insects) or the final molt to adulthood (in hemimetabolous insects).

2. Ecdysone

  • A steroid hormone produced by the prothoracic glands.
  • Ecdysone initiates molting at all stages of development.
  • The interaction between JH and ecdysone determines the type of molt (larval, pupal, or adult).

Mechanism: When JH levels are high along with ecdysone, the insect molts into another larval/nymphal stage. When JH levels are low and ecdysone is present, it triggers pupation (complete metamorphosis) or the final adult molt (incomplete metamorphosis).

Evolutionary Significance

Metamorphosis offers several evolutionary advantages:

  • Reduced Competition: Larvae and adults often occupy different niches, minimizing competition for resources.
  • Efficient Resource Utilization: Larval stages specialize in feeding and growth, while adults focus on reproduction and dispersal.
  • Increased Dispersal: The adult stage, often winged, facilitates dispersal to new habitats.

Comparison Table

Feature Complete Metamorphosis Incomplete Metamorphosis
Life Stages Egg, Larva, Pupa, Adult Egg, Nymph, Adult
Larva/Nymph Resemblance to Adult Very different Similar
Pupal Stage Present Absent
Ecological Niche Distinct larval and adult niches Overlapping niches

Conclusion

Metamorphosis is a defining characteristic of insect evolution, enabling remarkable adaptability and diversification. The interplay of juvenile hormone and ecdysone orchestrates this complex developmental process, allowing insects to exploit a wide range of ecological opportunities. Understanding the hormonal control and evolutionary advantages of metamorphosis is crucial for comprehending insect biology, pest management strategies, and the broader ecological dynamics of insect populations. Further research into the genetic and molecular mechanisms underlying metamorphosis continues to reveal the intricacies of insect development.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Ecdysis
The process of shedding the exoskeleton in arthropods, including insects, to allow for growth. It is also known as molting.
Corpora Allata
A pair of endocrine glands in insects that produce juvenile hormone, a key regulator of insect development and metamorphosis.

Key Statistics

Approximately 85% of all insect species undergo complete metamorphosis (Holometabolism).

Source: Chapman, R. F. (2013). *The Insects: Structure and Function*. Cambridge University Press.

Insect pests cause an estimated 20-25% of global crop losses annually.

Source: FAO (Food and Agriculture Organization of the United Nations), 2019.

Examples

Monarch Butterfly

The Monarch butterfly exemplifies complete metamorphosis. The larva (caterpillar) feeds voraciously on milkweed, accumulating resources for pupation. The pupa (chrysalis) undergoes a complete reorganization, and the adult butterfly emerges with wings for migration and reproduction.

Frequently Asked Questions

What is the role of the prothoracic gland?

The prothoracic gland is responsible for synthesizing and releasing ecdysone, the hormone that initiates molting at all stages of insect development. It's a crucial component in the hormonal control of metamorphosis.

Topics Covered

BiologyZoologyInsectsDevelopmental BiologyEndocrinology