UPSC MainsAGRICULTURE-PAPER-I201115 Marks150 Words
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Q17.

Define polyembryony. Discuss its significance in horticulture with special reference to mango and citrus.

How to Approach

This question requires a clear understanding of polyembryony and its horticultural significance. The approach should begin by defining polyembryony and then elaborating on its advantages in horticulture, specifically highlighting its application in mango and citrus cultivation. The answer should showcase how this phenomenon allows for clonal propagation and preservation of desirable traits. A concise and well-structured response demonstrating knowledge of horticultural practices is expected.

Model Answer

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Introduction

Polyembryony, a fascinating developmental anomaly, refers to the phenomenon where more than one embryo develops from a single fertilized ovule. This contrasts with the typical scenario where a single embryo forms. While initially considered a curiosity, polyembryony has gained considerable importance in horticulture, particularly for propagating superior varieties. The ability to generate multiple seedlings from a single seed offers a unique opportunity for clonal propagation, ensuring the preservation of desirable traits that are often lost in sexual reproduction. This is especially crucial for crops like mango and citrus, known for their complex genetic makeups and often exhibiting unpredictable inheritance patterns.

Defining Polyembryony

Polyembryony can be broadly classified into two types: adventive and semi-parthenocarpic. Adventive polyembryony arises spontaneously due to the division of the zygote or the formation of additional embryo sacs. Semi-parthenocarpic polyembryony, commonly found in mango, is a result of nucellar embryony where the nucellus (tissue surrounding the embryo sac) contributes to embryo formation. This is often linked to apomixis, a form of asexual seed production.

Significance in Horticulture

The significance of polyembryony in horticulture is primarily due to its potential for clonal propagation. This allows growers to reproduce plants with desirable traits like superior fruit quality, disease resistance, and high yield, which may not be consistently inherited through traditional seed propagation.

Mango (Mangifera indica)

In mango, polyembryony is predominantly of the semi-parthenocarpic type. It's a widely exploited characteristic in mango breeding and propagation. The nucellus, a nutritive tissue within the ovule, divides and forms multiple embryos. This means a single mango seed can produce several seedlings, all genetically identical to the mother plant. This is invaluable for preserving elite cultivars like 'Alphonso' and 'Kesar'.

Example: The 'Alphonso' mango variety, famed for its rich flavor, is frequently propagated using polyembryonic seeds to maintain its unique characteristics.

Citrus (Citrus spp.)

While less common than in mango, polyembryony does occur in citrus varieties. In citrus, it's typically adventive, arising from zygote division. The presence of multiple embryos allows for the selection of desirable seedlings, particularly those exhibiting rootstock characteristics for grafting. Grafting is a common practice in citrus cultivation to combine desirable fruit traits with robust root systems.

Case Study: The Citrus Research Institute in Florida has utilized polyembryony in breeding programs to select for rootstock varieties with resistance to citrus greening disease (Huanglongbing - HLB). By screening seedlings from polyembryonic seeds, researchers can identify plants with natural resistance and use them as rootstocks for grafting scions of commercially desirable fruit varieties.

Advantages of Utilizing Polyembryony in Horticulture

  • Clonal Propagation: Ensures the replication of desired traits.
  • Preservation of Elite Varieties: Protects unique cultivars from genetic degradation.
  • Simplified Propagation: Avoids complex grafting or tissue culture techniques.
  • Rootstock Selection (Citrus): Facilitates the selection of disease-resistant rootstocks.

Limitations

While advantageous, polyembryony isn’t without limitations. The resulting seedlings may exhibit variations, particularly in adventive polyembryony. Furthermore, the ratio of polyembryonic to monoembryonic seeds can vary, requiring careful selection.

Crop Type of Polyembryony Significance
Mango Semi-parthenocarpic Clonal propagation of elite cultivars
Citrus Adventive Selection of rootstocks for grafting

Conclusion

Polyembryony represents a significant advantage in horticulture, particularly for crops like mango and citrus, allowing for efficient clonal propagation and the preservation of valuable traits. While limitations exist, the benefits of utilizing this phenomenon far outweigh the drawbacks. Continued research into the mechanisms and control of polyembryony promises to further enhance its application in plant breeding and sustainable agricultural practices, ensuring the availability of superior varieties for future generations.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Apomixis
A form of asexual reproduction in plants where seeds are produced without fertilization. Polyembryony in mango is often linked to this process.
Nucellus
The central tissue of an ovule in flowering plants, which surrounds the embryo sac. In mango, the nucellus contributes to embryo formation in polyembryonic seeds.

Key Statistics

The percentage of polyembryonic seeds in mango varieties can vary from 10% to 50%, depending on the cultivar. (Source: Knowledge Cutoff)

Source: Knowledge Cutoff

Citrus greening disease (HLB) has caused billions of dollars in losses to the citrus industry worldwide.

Source: Knowledge Cutoff

Examples

‘Alphonso’ Mango Propagation

Farmers in Maharashtra, India, heavily rely on polyembryonic seeds to propagate ‘Alphonso’ mango trees, ensuring the continuation of this prized variety.

Frequently Asked Questions

Why is polyembryony not always desirable?

While beneficial for clonal propagation, polyembryony can sometimes lead to genetic variability among seedlings, potentially diluting the desired traits. Control over the process is therefore important.

Topics Covered

HorticultureBotanyPlant PropagationAsexual ReproductionMangoCitrus