UPSC MainsBOTANY-PAPER-II201220 Marks
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Q22.

Explain the process of anaerobic glucose catabolism and mention the mode of ATP generation during the process.

How to Approach

This question requires a detailed understanding of anaerobic respiration, specifically focusing on glucose catabolism. The answer should begin by defining anaerobic respiration and its significance. Then, it should meticulously explain the steps involved – glycolysis, fermentation (lactic acid and alcoholic), and the mechanisms of ATP generation in each stage. A comparative table highlighting ATP yield in aerobic vs. anaerobic respiration would be beneficial. Finally, the answer should emphasize the importance of this pathway in various organisms and physiological conditions.

Model Answer

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Introduction

Anaerobic glucose catabolism refers to the breakdown of glucose in the absence of oxygen. This metabolic pathway is crucial for organisms living in oxygen-deprived environments and also occurs in animal cells during intense physical activity when oxygen supply is limited. While less efficient than aerobic respiration, anaerobic pathways allow for the continued production of ATP, albeit in smaller quantities, enabling cells to maintain essential functions. Understanding this process is fundamental to comprehending cellular energy production and its implications in diverse biological systems. This process is a vital survival mechanism for many organisms and plays a significant role in various industrial applications like fermentation.

Anaerobic Glucose Catabolism: A Detailed Explanation

Anaerobic glucose catabolism can be broadly divided into two main phases: Glycolysis and Fermentation.

1. Glycolysis

Glycolysis is the initial step in both aerobic and anaerobic respiration. It occurs in the cytoplasm and involves the breakdown of one molecule of glucose (a 6-carbon sugar) into two molecules of pyruvate (a 3-carbon molecule). This process doesn't require oxygen and yields a net gain of 2 ATP molecules, 2 NADH molecules, and 2 pyruvate molecules.

  • Energy Investment Phase: 2 ATP molecules are used to phosphorylate glucose.
  • Energy Payoff Phase: 4 ATP molecules are produced, resulting in a net gain of 2 ATP.

2. Fermentation

Fermentation follows glycolysis when oxygen is absent. It regenerates NAD+ from NADH, which is essential for glycolysis to continue. There are two main types of fermentation:

a) Lactic Acid Fermentation

In lactic acid fermentation, pyruvate is reduced directly by NADH to form lactate as an end product. This process regenerates NAD+ allowing glycolysis to continue. This occurs in muscle cells during strenuous exercise and in certain bacteria used in yogurt production.

Reaction: Pyruvate + NADH → Lactate + NAD+

b) Alcoholic Fermentation

In alcoholic fermentation, pyruvate is first decarboxylated to acetaldehyde, releasing carbon dioxide. Acetaldehyde is then reduced by NADH to ethanol, regenerating NAD+. This process is carried out by yeasts and some bacteria and is used in brewing and baking.

Reactions:

  • Pyruvate → Acetaldehyde + CO2
  • Acetaldehyde + NADH → Ethanol + NAD+

ATP Generation During Anaerobic Glucose Catabolism

The primary mode of ATP generation during anaerobic glucose catabolism is substrate-level phosphorylation during glycolysis. This process directly generates ATP by transferring a phosphate group from a high-energy intermediate molecule to ADP. Fermentation itself does not directly produce ATP; its primary role is to regenerate NAD+ to sustain glycolysis.

The following table summarizes the ATP yield comparison between aerobic and anaerobic respiration:

Process ATP Yield (per glucose molecule) Oxygen Requirement
Aerobic Respiration ~36-38 ATP Required
Anaerobic Respiration (Glycolysis + Fermentation) 2 ATP Not Required

Significance and Applications

Anaerobic glucose catabolism is vital for several reasons:

  • Survival in Oxygen-Deficient Environments: Allows organisms to survive and function in the absence of oxygen.
  • Intense Exercise: Provides a rapid, albeit limited, source of ATP during strenuous activity.
  • Industrial Applications: Used in the production of various products like ethanol, lactic acid, yogurt, cheese, and bread.

Conclusion

In conclusion, anaerobic glucose catabolism is a crucial metabolic pathway that enables cells to generate ATP in the absence of oxygen. While less efficient than aerobic respiration, it provides a vital energy source for organisms in oxygen-limited environments and during periods of intense activity. The process, encompassing glycolysis and fermentation, relies on substrate-level phosphorylation for ATP production and highlights the adaptability of cellular metabolism to varying environmental conditions. Further research into optimizing anaerobic pathways could have significant implications for biofuel production and understanding metabolic diseases.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Glycolysis
The metabolic pathway that breaks down glucose into pyruvate, producing a small amount of ATP and NADH.
Substrate-level phosphorylation
A direct method of ATP synthesis where a phosphate group is transferred from a high-energy intermediate molecule to ADP.

Key Statistics

The human body can continue anaerobic metabolism for approximately 1-2 minutes during intense exercise before lactic acid buildup inhibits muscle function. (Source: Widmaier, E. P., Raff, H., & Strang, K. T. (2016). Vander's human physiology: The mechanisms of body function. McGraw-Hill Education.)

Source: Widmaier et al., 2016

Global ethanol production reached approximately 110 billion liters in 2022, largely driven by biofuel mandates and industrial applications. (Source: Renewable Fuels Association, 2023 - Knowledge Cutoff)

Source: Renewable Fuels Association, 2023

Examples

Yeast Fermentation in Bread Making

Yeast utilizes alcoholic fermentation to produce carbon dioxide, which causes bread dough to rise. The ethanol produced evaporates during baking.

Frequently Asked Questions

What happens to lactic acid produced during lactic acid fermentation?

Lactic acid is transported to the liver, where it can be converted back into glucose via the Cori cycle, or it can be oxidized for energy.

Topics Covered

BiologyBiochemistryRespirationMetabolismEnergy Production