UPSC MainsAGRICULTURE-PAPER-II201410 Marks150 Words
हिंदी में पढ़ें
Q20.

Show full reactions where (i) Alcohol dehydrogenase, (ii) Nitrate reductase, (iii) Glucokinase and (iv) Succinate dehydrogenase are involved, therewith mentioning respective co-factor, co-enzyme, prosthetic group etc., if so required.

How to Approach

This question requires a detailed understanding of enzymatic reactions crucial in plant metabolism. The approach should be to first briefly introduce each enzyme, then outline the full reaction it catalyzes, highlighting the substrates, products, and crucially, the cofactors/prosthetic groups involved. A tabular format may be beneficial for clarity. Focus on accuracy and completeness in detailing the biochemical pathways.

Model Answer

0 min read

Introduction

Enzymes are biological catalysts that accelerate biochemical reactions within living organisms. In plants, these enzymes play a pivotal role in various metabolic processes, including photosynthesis, respiration, and nitrogen metabolism. The enzymes mentioned – Alcohol dehydrogenase, Nitrate reductase, Glucokinase, and Succinate dehydrogenase – are vital for these processes. Understanding their mechanisms and the cofactors they utilize is fundamental to comprehending plant physiology. This response will detail each enzyme's reaction, including the necessary cofactors, coenzymes, and prosthetic groups.

Alcohol Dehydrogenase (ADH)

Alcohol dehydrogenase catalyzes the reversible oxidation of alcohols to aldehydes or ketones. It's crucial in ethanol metabolism and detoxification.

Reaction: CH3CH2OH + NAD+ ⇌ CH3CHO + NADH + H+

Cofactor/Coenzyme: NAD+ (Nicotinamide Adenine Dinucleotide) acts as the coenzyme, accepting electrons and hydrogen ions during the oxidation process. It's reduced to NADH.

Nitrate Reductase (NR)

Nitrate reductase is essential for nitrogen assimilation in plants, converting nitrate to nitrite, a necessary step for incorporating nitrogen into amino acids and proteins.

Reaction: NO3- + NADH + H+ ⇌ NO2- + NAD+ + H2O

Cofactor/Prosthetic Group: NR is a complex enzyme with multiple components. It requires FAD (Flavin Adenine Dinucleotide) as a prosthetic group bound to the catalytic site, and a heme group (iron-containing porphyrin ring) for electron transfer. NADH acts as the reducing agent, a coenzyme.

Glucokinase (GK)

Glucokinase, also known as hexokinase, catalyzes the phosphorylation of glucose to glucose-6-phosphate. This is a key step in glucose metabolism, especially in photosynthetic tissues.

Reaction: Glucose + ATP ⇌ Glucose-6-Phosphate + ADP

Cofactor/Coenzyme: ATP (Adenosine Triphosphate) is the substrate providing the phosphate group. Magnesium ions (Mg2+) are essential as cofactors, facilitating ATP binding and phosphate transfer.

Succinate Dehydrogenase (SDH)

Succinate dehydrogenase is a vital enzyme complex (Complex II) in both the citric acid cycle (Krebs cycle) and the electron transport chain. It catalyzes the oxidation of succinate to fumarate.

Reaction: Succinate + FAD ⇌ Fumarate + FADH2

Cofactor/Prosthetic Group: SDH contains FAD (Flavin Adenine Dinucleotide) as a prosthetic group, which is reduced to FADH2 during the oxidation of succinate. It also contains heme groups (iron-containing porphyrin rings) involved in electron transfer to ubiquinone.

Enzyme Reaction Cofactor/Coenzyme/Prosthetic Group
Alcohol Dehydrogenase CH3CH2OH + NAD+ ⇌ CH3CHO + NADH + H+ NAD+
Nitrate Reductase NO3- + NADH + H+ ⇌ NO2- + NAD+ + H2O FAD, Heme, NADH
Glucokinase Glucose + ATP ⇌ Glucose-6-Phosphate + ADP ATP, Mg2+
Succinate Dehydrogenase Succinate + FAD ⇌ Fumarate + FADH2 FAD, Heme

Conclusion

In conclusion, the enzymes Alcohol dehydrogenase, Nitrate reductase, Glucokinase, and Succinate dehydrogenase are crucial components of plant metabolism, each playing a distinct role in vital processes. Their activity relies on specific cofactors, coenzymes, and prosthetic groups, which are essential for their catalytic function. Understanding these enzymatic reactions and their associated factors is critical for a comprehensive understanding of plant physiology and biochemistry.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Coenzyme
Organic non-protein molecules that bind to enzymes and are required for their catalytic activity. They are often derived from vitamins.
Prosthetic Group
A non-protein chemical compound that is tightly bound to an enzyme and is essential for its activity. Unlike coenzymes, they are usually permanently attached to the enzyme.

Key Statistics

Nitrate reductase activity is significantly influenced by light and carbohydrate availability, with optimal activity observed during periods of photosynthesis.

Source: Plant Physiology, 1987; 84(4): 655-661

Glucokinase plays a critical role in maintaining glucose homeostasis, preventing excessive glucose accumulation in photosynthetic tissues. Its activity is tightly regulated by ATP levels.

Examples

Ethanol Metabolism in Fermented Beverages

Alcohol dehydrogenase is crucial in the production of alcoholic beverages like beer and wine, where it converts sugars into ethanol.

Nitrogen Fixation in Legumes

Nitrate reductase's activity is essential for the efficient assimilation of nitrogen, a limiting nutrient in many ecosystems. Legumes benefit from symbiotic nitrogen fixation, increasing nitrate availability.

Frequently Asked Questions

What is the difference between a coenzyme and a prosthetic group?

Coenzymes are often transiently associated with the enzyme, while prosthetic groups are tightly bound and generally remain permanently attached.

How is nitrate reductase activity regulated in plants?

Nitrate reductase activity is regulated by light, carbohydrate availability, and the presence of inorganic phosphate.

Topics Covered

BotanyBiochemistryPlant PhysiologyEnzymologyMetabolism