UPSC MainsAGRICULTURE-PAPER-I201720 Marks
हिंदी में पढ़ें
Q9.

Define heterosis. How is heterosis being exploited in the development of hybrids?

How to Approach

This question requires a clear understanding of plant breeding principles. The approach should begin by defining heterosis and explaining its genetic basis. Then, elaborate on how breeders exploit heterosis through hybrid development, detailing the process and providing examples of successful hybrids in Indian agriculture. Finally, discuss the limitations and challenges associated with utilizing heterosis. A structured approach with clear headings and bullet points is crucial for a comprehensive answer.

Model Answer

0 min read

Introduction

Heterosis, also known as hybrid vigor, is a phenomenon observed in hybrid offspring that exhibit superior traits compared to their parents. This phenomenon has been a cornerstone of agricultural advancements, particularly in the development of high-yielding crop varieties. The Green Revolution in India, for example, heavily relied on hybrid seeds to increase food production. The concept was first scientifically explored by George Harrison Shull in the early 20th century. Understanding the genetic mechanisms behind heterosis and its practical application is vital for ensuring food security and improving agricultural productivity in a changing climate.

Defining Heterosis

Heterosis (H) is the superior performance of a hybrid (F1 generation) over the average performance of its inbred parental lines. It manifests as increased vigor, growth rate, yield, disease resistance, and other desirable traits. The term "hybrid vigor" is often used interchangeably with heterosis. It’s not simply a matter of the average of the parents being exceeded; it's often a dramatic and significant improvement.

Genetic Basis of Heterosis

The genetic basis of heterosis is complex and not fully understood. Several hypotheses have been proposed, including:

  • Dominance Hypothesis: This is the most widely accepted explanation. It suggests that dominant alleles present in one parent mask the recessive, less favorable alleles in the other parent, resulting in a superior phenotype in the hybrid.
  • Overdominance Hypothesis: This hypothesis proposes that the heterozygous condition at specific loci is advantageous compared to either homozygous condition. The interaction between different alleles at a locus leads to a more favorable phenotype.
  • Epistasis: Gene interactions (epistasis) can also contribute to heterosis. The effect of one gene is masked or modified by another.
  • Additive Effects: While heterosis is primarily driven by non-additive genetic effects (dominance and epistasis), additive gene effects also play a minor role.

Exploiting Heterosis in Hybrid Development

Breeders exploit heterosis through a carefully planned breeding process. The typical steps involved are:

  • Inbreeding: Repeated self-pollination of plants to create homozygous inbred lines. This process exposes recessive alleles and allows for selection of desirable traits. Inbreeding depression is a common challenge at this stage.
  • Cross-Pollination: Crossing two different inbred lines to generate the F1 hybrid. The F1 generation exhibits heterosis.
  • Selection of Parental Lines: Careful selection of parental lines is crucial. Lines should be genetically diverse and possess complementary desirable traits.
  • Hybrid Evaluation: Rigorous testing of the hybrid under various environmental conditions to ensure consistent performance.

Examples of Hybrid Exploitation in Indian Agriculture

India has been a global leader in the adoption of hybrid technology. Some prominent examples include:

  • Hybrid Maize: India is one of the largest producers of hybrid maize globally. Hybrid maize varieties offer significantly higher yields compared to traditional varieties. The Indian Council of Agricultural Research (ICAR) has been instrumental in developing and disseminating hybrid maize seeds.
  • Hybrid Cotton: The introduction of Bt cotton hybrids (genetically modified to resist bollworms) has revolutionized cotton production in India. While controversial, these hybrids have increased yields and reduced pesticide use.
  • Hybrid Rice: Hybrid rice varieties have gained popularity due to their higher yields and drought tolerance. Companies like Bayer (Monsanto) have played a significant role in hybrid rice development.
  • Hybrid Vegetables: Hybrid varieties of vegetables like tomato, chili, and eggplant are widely cultivated to enhance productivity and quality.

Challenges and Limitations

Despite its benefits, utilizing heterosis faces certain challenges:

  • Inbreeding Depression: The process of creating inbred lines leads to inbreeding depression, reducing vigor and yield.
  • Cost and Complexity: Hybrid seed production is more expensive and complex than producing open-pollinated varieties.
  • Hybrid Seed Dependency: Farmers are dependent on seed companies for hybrid seeds, as the F2 generation (next generation after the hybrid) typically loses the heterotic advantage. This raises concerns about farmer dependency and cost.
  • Genetic Erosion: Focus on a few high-yielding hybrids can lead to genetic erosion of traditional varieties.
  • Environmental Concerns: The widespread use of hybrid seeds, especially genetically modified hybrids, raises environmental concerns regarding biodiversity and pesticide resistance.
Aspect Inbred Lines Hybrid (F1)
Yield Lower Higher
Vigor Reduced (Inbreeding Depression) Enhanced (Heterosis)
Genetic Uniformity High Lower
Seed Production Cost Lower Higher

Conclusion

Heterosis remains a vital tool for enhancing agricultural productivity, particularly in a world facing increasing food security challenges. While the process of developing and utilizing hybrids presents complexities and limitations, ongoing research focusing on genetic diversity, improved inbred lines, and sustainable practices can further maximize the benefits of heterosis. Investing in public sector research and ensuring farmer access to quality hybrid seeds at affordable prices is crucial for realizing the full potential of this phenomenon and promoting a resilient and sustainable agricultural system.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Inbreeding Depression
The reduction in vigor and yield observed in self-pollinated plants due to the expression of deleterious recessive alleles.
Epistasis
The interaction between genes where the effect of one gene is masked or modified by another gene.

Key Statistics

Hybrid maize yields in India are typically 30-50% higher than those of conventional maize varieties. (Source: ICAR – Knowledge Cutoff)

Source: ICAR

Bt cotton hybrids account for approximately 95% of cotton cultivation in India. (Source: Ministry of Agriculture & Farmers Welfare – Knowledge Cutoff)

Source: MoAFW

Frequently Asked Questions

Why does the F2 generation of hybrids lose the heterotic advantage?

The F2 generation results from self-pollination or cross-pollination within the F1 generation. This segregation of genes breaks down the genetic combination that created the heterosis in the F1 generation.

Topics Covered

AgriculturePlant BreedingGeneticsHeterosisHybrid SeedsPlant BreedingGenetics