UPSC MainsAGRICULTURE-PAPER-II201720 Marks
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Q12.

What are mutations? Indicate the use of induced mutation in varietal development.

How to Approach

This question requires a clear understanding of mutation and its application in plant breeding. The approach should begin by defining mutations and their types, followed by an explanation of induced mutations and their significance in varietal development. The response should include examples of successful induced mutations leading to improved crop varieties. A structured approach with clear headings and subheadings is essential for a comprehensive answer.

Model Answer

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Introduction

Mutations are the driving force behind genetic diversity and evolution, playing a pivotal role in agriculture. They represent alterations in the DNA sequence, which can arise spontaneously or be induced artificially. The Green Revolution, which dramatically increased food production in the mid-20th century, heavily relied on the principles of plant breeding, including the exploitation of mutations. Recent advancements in CRISPR-Cas9 technology are further revolutionizing the landscape of plant breeding, offering unprecedented precision in inducing desired mutations. This response will explore the nature of mutations and detail how induced mutations are employed in the development of new and improved crop varieties.

What are Mutations?

Mutations are changes in the nucleotide sequence of an organism's genome. These changes can occur in any part of the genome, including coding and non-coding regions. They are the ultimate source of genetic variation, providing the raw material for natural selection and artificial selection.

Types of Mutations

  • Spontaneous Mutations: These occur naturally due to errors during DNA replication, repair, or recombination. They are relatively infrequent.
  • Induced Mutations: These are caused by exposure to external agents like radiation (UV, X-rays, gamma rays) or chemical mutagens. These are often used in plant breeding.

Mutations can be classified based on their effect:

  • Point Mutations: Changes in a single nucleotide (e.g., substitution, insertion, deletion).
  • Frameshift Mutations: Insertions or deletions that alter the reading frame of the genetic code.
  • Chromosomal Mutations: Large-scale changes in chromosome structure (e.g., deletion, duplication, inversion, translocation).

Induced Mutations in Varietal Development

Induced mutations are a powerful tool in plant breeding. By exposing seeds or plant tissues to mutagens, breeders can generate a vast number of new genetic combinations. The process involves screening these mutants for desirable traits.

Mechanism of Induced Mutations

Mutagens cause damage to DNA, leading to base modifications, strand breaks, and chromosomal aberrations. These damages are then repaired by the plant's DNA repair mechanisms, but these repairs are often imperfect, resulting in permanent changes in the DNA sequence – the induced mutations.

Process of Induced Mutation Breeding

  1. Mutagenesis: Seeds or plant tissues are exposed to mutagens (e.g., ethyl methanesulfonate (EMS), sodium azide). EMS is commonly used to induce point mutations.
  2. Selection: The M1 (first mutant) generation is allowed to self-pollinate, and seeds are harvested. The M2 generation is screened for desired traits.
  3. Stabilization: Mutants with desirable traits are selected and self-pollinated for several generations to achieve genetic stability (usually M3 to M5 generations).
  4. Release: The stabilized mutant lines are evaluated for yield, quality, and disease resistance before release to farmers.

Examples of Successful Induced Mutations

Crop Trait Improved Mutagen Used Year
Rice Semi-dwarfism, disease resistance Gamma rays 1960s
Wheat Improved protein content, rust resistance EMS 1970s
Groundnut High oil content, disease resistance Gamma rays 1980s
Rice (Shikar – a variety) High grain yield, resistance to blast disease EMS 1990s

The development of Shikar rice is a prime example of successful induced mutagenesis. It showcases how targeted mutations can lead to substantial improvements in crop yield and disease resistance. The National Plant Genetic Stock Repository (NPGR) maintains a vast collection of induced mutant lines for future use.

Challenges and Future Directions

  • Randomness: Induced mutations are largely random, requiring extensive screening.
  • Undesirable Mutations: Mutagenesis can also generate undesirable traits, requiring careful selection.
  • Genetic Instability: Some induced mutants may exhibit genetic instability.

Modern techniques like CRISPR-Cas9 gene editing offer greater precision in inducing mutations, allowing breeders to target specific genes and minimize the generation of undesirable traits. This technology is poised to revolutionize plant breeding, leading to faster and more efficient development of improved crop varieties.

Conclusion

In conclusion, mutations, both spontaneous and induced, are fundamental to genetic diversity and crop improvement. Induced mutations, while initially a random process, have proven invaluable in developing improved crop varieties, as exemplified by the success of Shikar rice. The advent of precision gene-editing technologies like CRISPR-Cas9 promises to refine this process further, offering targeted and efficient solutions for addressing the challenges of food security and climate change. Continued research and responsible application of these technologies are crucial for sustainable agricultural development.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Mutagenesis
The process of inducing mutations in an organism's DNA using physical or chemical agents.
NPGR
National Plant Genetic Stock Repository - A repository maintaining genetic resources, including induced mutants, for research and breeding purposes.

Key Statistics

Approximately 30% of crop varieties released worldwide have been developed using induced mutation techniques. (Source: FAO, Knowledge Cutoff)

Source: FAO

EMS is estimated to induce around 1 mutation per 10<sup>6</sup> base pairs in the genome. (Source: Knowledge Cutoff)

Source: Knowledge Cutoff

Examples

Shikar Rice

A high-yielding, blast-resistant rice variety developed through EMS mutagenesis, significantly contributing to rice production in India.

Frequently Asked Questions

What is the difference between spontaneous and induced mutations?

Spontaneous mutations occur naturally due to errors in DNA replication or repair, while induced mutations are caused by external factors like radiation or chemicals.

Topics Covered

AgricultureBotanyPlant BreedingMutationsPlant BreedingGeneticsVarietal Development