UPSC MainsAGRICULTURE-PAPER-I201810 Marks150 Words
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Q16.

Explain C3, C4 and CAM mechanism in plants and their importance in changing climatic conditions.

How to Approach

This question requires a clear understanding of C3, C4, and CAM photosynthetic pathways. The approach should be to first define each pathway, detailing the biochemical mechanisms involved. Subsequently, highlight the advantages and disadvantages of each, and finally, discuss their relevance in the context of changing climatic conditions like increased temperatures and CO2 levels. Structure the answer using a tabular format for comparison and concise explanations for each mechanism.

Model Answer

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Introduction

Photosynthesis, the cornerstone of life on Earth, isn't a monolithic process. Plants have evolved diverse strategies to capture sunlight and convert it into energy. These strategies are broadly categorized into C3, C4, and CAM photosynthesis, differing significantly in their carbon fixation mechanisms. The ongoing climate crisis, characterized by rising temperatures and altered precipitation patterns, is placing immense pressure on plant physiology. Understanding these photosynthetic pathways and their adaptability is crucial for ensuring food security and ecosystem resilience in a changing world. This answer will elucidate these mechanisms and their implications in the face of climate change.

C3 Photosynthesis

C3 photosynthesis is the most common pathway, utilized by approximately 85% of plant species. It involves the initial fixation of CO2 by RuBisCO (Ribulose-1,5-bisphosphate carboxylase/oxygenase) into a 3-carbon compound, 3-phosphoglycerate (3-PGA). This process occurs in the mesophyll cells. However, RuBisCO also binds to oxygen, leading to photorespiration, especially at high temperatures and low CO2 concentrations.

C4 Photosynthesis

C4 photosynthesis is an adaptation to hot, arid environments. It involves initial CO2 fixation by PEP carboxylase (PEPcase) in mesophyll cells, forming a 4-carbon compound (oxaloacetate). This is then transported to bundle sheath cells, where it’s decarboxylated, releasing CO2 that is then fixed by RuBisCO. This spatial separation minimizes photorespiration. Examples include maize, sugarcane, and sorghum.

CAM (Crassulacean Acid Metabolism) Photosynthesis

CAM photosynthesis is another adaptation to arid conditions, primarily found in succulent plants like cacti and pineapple. It temporally separates CO2 fixation. At night, stomata open, allowing CO2 to be fixed by PEPcase into a 4-carbon acid. During the day, stomata close to conserve water, and the stored acid is decarboxylated, releasing CO2 for fixation by RuBisCO. This temporal separation minimizes water loss and photorespiration.

Comparison of Photosynthetic Pathways

Feature C3 C4 CAM
Initial CO2 Fixation Enzyme RuBisCO PEPcase PEPcase (night), RuBisCO (day)
Cell Type for Initial Fixation Mesophyll Mesophyll Mesophyll (night), Bundle Sheath (day)
Photorespiration High Low Low
Water Use Efficiency Low High Very High
Temperature Optimum 15-25°C 30-40°C 20-35°C
Examples Rice, Wheat, Soybeans Maize, Sugarcane, Sorghum Cacti, Pineapple, Orchids

Importance in Changing Climatic Conditions

Rising temperatures and increasing CO2 concentrations are impacting plant physiology. C3 plants suffer from increased photorespiration at higher temperatures. C4 and CAM plants, with their mechanisms to minimize photorespiration and conserve water, are better adapted to these conditions. The increasing CO2 levels initially favor C3 plants, but the advantage diminishes with increasing temperatures. There's ongoing research into engineering C3 plants to exhibit C4-like traits (C4 rice project) to improve yields under stressful conditions.

Conclusion

In conclusion, C3, C4, and CAM photosynthesis represent distinct evolutionary adaptations to varying environmental conditions. As climate change intensifies, understanding these pathways and their limitations is critical for ensuring sustainable agriculture and preserving biodiversity. While C4 and CAM plants hold inherent advantages in stressful environments, ongoing research focused on improving the efficiency of C3 photosynthesis, particularly through genetic engineering, offers a crucial pathway to safeguard global food security in a rapidly changing world. Future strategies must integrate these adaptations with sustainable agricultural practices.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Topics Covered

BotanyClimate ChangePhotosynthesisPlant PhysiologyAdaptation