UPSC MainsAGRICULTURE-PAPER-II201810 Marks150 Words
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Q16.

Explain C3, C4 and CAM mechanism in plants and their importance in changing climatic conditions.

How to Approach

This question requires a concise explanation of three photosynthetic pathways – C3, C4, and CAM. The approach should be to define each pathway, outline their mechanisms, and then discuss their relevance in the context of climate change, specifically focusing on water use efficiency and adaptation strategies. A table comparing the three pathways would be beneficial for clarity. The answer must be structured to address all aspects of the question within the word limit.

Model Answer

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Introduction

Photosynthesis, the cornerstone of life on Earth, is the process by which plants convert light energy into chemical energy. While the basic principle remains the same, different plant species have evolved diverse photosynthetic mechanisms to thrive in varied environments. Among these are C3, C4, and CAM photosynthesis. These pathways differ primarily in the initial carbon fixation process and are increasingly important to understand in the face of accelerating climate change and its impacts on water availability and temperature. This answer will explain each mechanism and their significance in a changing climate.

C3 Photosynthesis

C3 photosynthesis is the most common pathway, utilized by approximately 85% of plant species, including wheat, rice, and soybeans. In this process, carbon dioxide is directly fixed by the enzyme RuBisCO (Ribulose-1,5-bisphosphate carboxylase/oxygenase) to form a 3-carbon compound – 3-phosphoglycerate (3-PGA). This process occurs in the mesophyll cells. However, RuBisCO also binds to oxygen, leading to photorespiration, which reduces photosynthetic efficiency, particularly in hot and dry conditions.

C4 Photosynthesis

C4 photosynthesis is an adaptation to hot and dry climates. Plants like maize, sorghum, and sugarcane utilize this pathway. In C4 plants, CO2 is initially fixed in mesophyll cells by the enzyme PEP carboxylase (PEPcase), forming a 4-carbon compound (oxaloacetate). This is then transported to bundle sheath cells, where it's decarboxylated, releasing CO2 which then enters the Calvin cycle. This spatial separation minimizes photorespiration by concentrating CO2 around RuBisCO. C4 plants exhibit higher water use efficiency.

CAM Photosynthesis

CAM (Crassulacean Acid Metabolism) photosynthesis is another adaptation to arid conditions, found in plants like cacti, succulents, and pineapple. CAM plants open their stomata at night to fix CO2 using PEPcase, storing it as malic acid in vacuoles. During the day, stomata close to conserve water, and the stored malic acid is decarboxylated, releasing CO2 for the Calvin cycle. This temporal separation of carbon fixation and the Calvin cycle allows CAM plants to survive in extremely dry environments.

Comparison Table

Feature C3 C4 CAM
Initial CO2 Fixation RuBisCO PEPcase PEPcase (night)
Cell Type Mesophyll Mesophyll & Bundle Sheath Mesophyll
Photorespiration High Low Low
Water Use Efficiency Low High Very High
Habitat Temperate, Moist Hot, Dry Arid

Importance in Changing Climatic Conditions

As climate change intensifies, with increased temperatures and water scarcity, understanding these photosynthetic pathways becomes crucial. C4 and CAM plants demonstrate superior water use efficiency compared to C3 plants. This makes them valuable for breeding programs aimed at developing drought-resistant crops. Furthermore, understanding the genetic mechanisms underlying C4 and CAM photosynthesis can help in engineering C3 plants to be more resilient to climate change impacts. The global average crop yield loss due to drought is estimated to be around 8% (FAO, 2019). Utilizing these pathways can mitigate these losses.

Conclusion

In conclusion, C3, C4, and CAM photosynthesis represent distinct adaptations to varying environmental conditions. As climate change exacerbates water scarcity and increases temperatures, understanding and leveraging the advantages of C4 and CAM pathways through breeding and genetic engineering is vital for ensuring food security and sustainable agriculture. Further research into these pathways holds significant promise for developing climate-resilient crops and mitigating the impacts of a changing climate on global food production.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Photorespiration
A metabolic pathway that occurs when RuBisCO binds to oxygen instead of carbon dioxide, reducing photosynthetic efficiency.
RuBisCO
Ribulose-1,5-bisphosphate carboxylase/oxygenase, an enzyme crucial for carbon fixation in photosynthesis.

Key Statistics

Approximately 85% of plant species on Earth utilize C3 photosynthesis.

Source: Raven, J. A. (1997). Photosynthesis in terrestrial plants.

CAM plants can have a water use efficiency that is 5-10 times greater than C3 plants.

Source: Nobel, P. S. (1988). Environmental physiology of wild plants.

Examples

Maize (C4 plant)

Maize, a staple food crop, utilizes C4 photosynthesis, exhibiting higher photosynthetic rates and water use efficiency compared to C3 crops like wheat.

Frequently Asked Questions

Why are C4 and CAM plants more efficient in hot, dry climates?

C4 and CAM plants minimize photorespiration and conserve water by spatially or temporally separating the initial carbon fixation and the Calvin cycle.

Topics Covered

BotanyClimate ChangePhotosynthesisPlant PhysiologyAdaptation