UPSC MainsAGRICULTURE-PAPER-II201810 Marks150 Words
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Q3.

What do you mean by heterosis? Describe the physiological bases of heterosis.

How to Approach

This question requires a clear understanding of heterosis and its underlying physiological mechanisms. The approach should be to first define heterosis, then explain its significance in crop improvement. Following this, a detailed discussion of the genetic and physiological bases of heterosis, including dominance and overdominance hypotheses, should be provided. Finally, the practical implications of heterosis in agriculture should be briefly mentioned. Structure: Definition, Significance, Genetic Basis, Physiological Basis, Practical Applications.

Model Answer

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Introduction

Heterosis, also known as hybrid vigor, is a phenomenon observed in hybrid offspring exhibiting superior traits compared to their parents. This phenomenon is particularly valuable in agriculture, leading to significant improvements in crop yield and quality. The Green Revolution, for instance, heavily relied on hybrid varieties exhibiting heterosis. Understanding heterosis is crucial for developing high-yielding crop varieties and ensuring food security. The concept gained prominence in the early 20th century with the work of George Harrison Shull and Edward Murray East.

What is Heterosis?

Heterosis refers to the superior performance of hybrid offspring compared to their inbred parents. This can manifest as increased yield, larger size, improved disease resistance, or other desirable traits. It's a cornerstone of modern hybrid crop breeding, particularly in maize, rice, and sugarcane.

Significance of Heterosis in Agriculture

  • Increased Yield: Hybrid varieties often exhibit significantly higher yields than pure lines.
  • Improved Quality: Heterosis can enhance traits like grain size, protein content, and nutritional value.
  • Disease Resistance: Hybrid vigor can confer resistance to various pests and diseases.
  • Adaptability: Hybrids can often adapt better to diverse environmental conditions.

Genetic Basis of Heterosis

Several hypotheses attempt to explain the genetic basis of heterosis:

1. Dominance Hypothesis:

This hypothesis suggests that heterosis arises due to the masking of deleterious recessive alleles in the inbred parents. When these parents are crossed, the hybrid offspring possess a dominant allele for each trait, masking the effect of the recessive alleles. While this can explain some aspects of heterosis, it doesn't fully account for all observed effects.

2. Overdominance Hypothesis:

This hypothesis proposes that heterozygotes (hybrid offspring) have a selective advantage over both homozygous parents due to the interaction of different alleles at the same locus. The combined effect of alleles leads to superior performance. This is considered a primary driver of heterosis for many traits.

3. Epistasis:

Epistasis refers to the interaction between genes at different loci. This interaction can contribute to heterosis by creating synergistic effects, leading to enhanced traits in the hybrid offspring.

Physiological Bases of Heterosis

The genetic changes resulting in heterosis translate into physiological and biochemical alterations:

  • Increased Photosynthesis: Hybrid plants often exhibit higher photosynthetic rates due to improved leaf area, chlorophyll content, or carbon dioxide assimilation.
  • Enhanced Nutrient Uptake: Heterosis can lead to improved root development and nutrient absorption, ensuring adequate supply for growth.
  • Improved Water Use Efficiency: Hybrid plants may demonstrate better water use efficiency, crucial for survival in water-stressed environments.
  • Altered Hormonal Balance: Changes in plant hormone levels (e.g., gibberellins, auxins) can contribute to increased growth and development in hybrid offspring.
  • Metabolic Efficiency: Heterosis can lead to more efficient metabolic processes, reducing energy expenditure and improving overall performance.

Example: In maize, hybrid varieties bred from inbred lines exhibit significantly higher grain yields (often 15-20% higher) compared to the inbred lines themselves, largely due to the overdominance effect at multiple loci.

Practical Applications

Heterosis is widely exploited in crop breeding programs, particularly for self-pollinating crops like maize, rice, wheat, and sorghum. The development of hybrid seeds is a multi-billion dollar industry globally. The National Seeds Corporation (NSC) plays a crucial role in promoting hybrid seed production and distribution in India.

Case Study: Maize Hybrid Development in India

The introduction of hybrid maize in India during the 1960s revolutionized maize production. Initially, the hybrid varieties were expensive and accessible only to larger farmers. However, with advancements in breeding techniques and increased availability of affordable hybrid seeds, maize production has significantly increased, contributing to food security and farmer income. The case study highlights the power of heterosis in transforming agricultural productivity.

Conclusion

In conclusion, heterosis is a vital phenomenon in agriculture, leading to substantial improvements in crop yield and quality. Driven by a combination of dominance, overdominance, and epistatic interactions, it results in significant physiological changes that enhance plant performance. Continued research and development of hybrid varieties remain crucial for ensuring food security and improving agricultural productivity in the face of climate change and increasing population demands.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Inbred Lines
Inbred lines are the result of repeated self-pollination, leading to reduced genetic diversity and homozygosity. They are essential for creating hybrid varieties exhibiting heterosis.
Epistasis
Epistasis is a phenomenon where the effect of one gene is masked or modified by the action of one or more other genes.

Key Statistics

Hybrid maize yields in India are, on average, 30-50% higher than those of traditional varieties (Source: ICAR-Indian Agricultural Research Institute, knowledge cutoff).

Source: ICAR-IARI

The global hybrid seed market is estimated to be worth over $13 billion (Source: Industry reports, knowledge cutoff).

Source: Industry Reports

Examples

Sugarcane Hybrid Development

Similar to maize, sugarcane breeders utilize heterosis to develop high-yielding hybrid varieties. These hybrids exhibit increased sucrose content and improved stalk thickness, contributing to higher sugar production.

Frequently Asked Questions

Why are hybrid seeds more expensive than open-pollinated seeds?

Hybrid seed production is a complex and resource-intensive process, involving multiple generations of breeding and careful quality control. This higher production cost is reflected in the price of hybrid seeds.

Topics Covered

AgricultureGeneticsPlant BreedingHybridizationGenetics