UPSC MainsAGRICULTURE-PAPER-II202115 Marks
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Q13.

What do you understand by centre of origin ? Discuss the law of homologous series of variation.

How to Approach

This question requires a nuanced understanding of plant evolution and genetics. The approach should be to first define "centre of origin" and its significance in crop improvement. Then, explain the Law of Homologous Series with clear examples, highlighting its importance in understanding genetic variation and plant breeding. A structured answer with clear headings and subheadings, coupled with relevant examples, will demonstrate a comprehensive understanding of the topic. Diagrams (if permitted) could further enhance clarity.

Model Answer

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Introduction

The origin and distribution of plant species are fundamental to understanding biodiversity and agricultural practices. The concept of “centre of origin” is crucial in identifying regions where wild relatives of crop plants evolved, retaining valuable genetic resources. These regions often represent hotspots of genetic diversity, vital for improving crop resilience and productivity. The Law of Homologous Series, proposed by August Wilhelm von Hofmann, provides a framework for understanding predictable variations in chemical and biological properties within a group of related compounds, a principle that finds application in plant genetics and breeding. This answer will explore both these concepts in detail, demonstrating their interconnectedness in the realm of plant evolution.

What is a Centre of Origin?

A "centre of origin" refers to the geographic region where a plant species first evolved or originated. It's the area where the wild relatives of a crop plant are most abundant and possess the greatest genetic diversity. Unlike centres of adaptation or cultivation, which are regions where plants have been modified by human selection, centres of origin represent the cradle of a species’ existence.

The concept was popularized by Nikolai Vavilov, a Russian botanist and geneticist, who established twelve primary centres of origin of cultivated plants based on his extensive expeditions and observations. These centres are:

  • Near East (Fertile Crescent)
  • Ethiopia
  • South Asia
  • East Asia
  • Americas (South and Meso)
  • Australasia
  • Africa
  • Mediterranean region
  • Indo-Malayasia
  • North America
  • Central America
  • South America

Vavilov’s work, though later challenged by molecular data, remains a cornerstone in understanding crop diversity. The significance of these centres lies in their potential to provide genes for disease resistance, drought tolerance, and improved yield.

The Law of Homologous Series

The Law of Homologous Series, developed by August Wilhelm von Hofmann in the 19th century, initially described predictable variations in the properties of organic compounds. It states that a group of chemical compounds with similar structures (homologous series) exhibit a gradual and predictable change in their physical and chemical properties as the molecular weight increases. This principle, while initially chemical, has been adapted to understand genetic variation in plants.

Applying the Law to Plant Genetics

In plant genetics, the homologous series concept is used to understand how variations arise within a group of related plants. For example, consider the series of wheat species (e.g., *Triticum monococcum*, *Triticum dicoccum*, *Triticum aestivum*). Each species shares a similar basic structure, but differs in chromosome number, grain size, and yield potential. These differences arise due to mutations and genetic recombination over time, following a somewhat predictable pattern.

Key Aspects of the Law in Plant Genetics

  • Structural Similarity: Plants within a homologous series share a common ancestral origin and possess similar fundamental structures.
  • Gradual Variation: Traits change progressively rather than abruptly.
  • Predictability: The changes in traits can often be predicted based on the underlying genetic differences.
  • Genetic Basis: These variations are often attributed to mutations in homologous genes.

Example: The Tomato Homologous Series

The wild tomato (*Solanum pimpinellifolium*) has been a source of numerous genes for improving cultivated tomatoes. Successive crosses and selections have led to a series of tomato varieties exhibiting gradual changes in fruit size, shape, and color. These changes, while influenced by selection, are rooted in the underlying genetic homology between the wild and cultivated forms.

Relationship between Centre of Origin and Homologous Series

The concept of centres of origin is intrinsically linked to the Law of Homologous Series. Centres of origin are the reservoirs of genetic diversity – the raw material upon which natural selection and subsequent artificial selection act. The variations observed within a homologous series within a plant species are often most pronounced and diverse within its centre of origin. Exploring these centres allows breeders to identify and utilize these variations for crop improvement.

Challenges and Modern Perspectives

While Vavilov’s centres remain important, modern molecular techniques have revealed a more complex picture. Genetic introgression (the transfer of genes from one species to another) and ancient hybridization events have blurred the lines between centres. Nevertheless, the principle of centres of origin as regions of high genetic diversity remains valid.

Concept Definition Significance
Centre of Origin Geographic region where a plant species originated. Source of genetic diversity for crop improvement.
Law of Homologous Series Predictable variation in traits within a group of related plants. Provides framework for understanding genetic variation and plant breeding.

Conclusion

In conclusion, understanding the concept of centres of origin and the Law of Homologous Series is crucial for appreciating the evolution and genetic diversity of plants. Vavilov’s centres provide vital gene pools for crop improvement, while the Law of Homologous Series helps to interpret the patterns of genetic variation within these pools. As genetic technologies advance, a more refined understanding of these concepts, coupled with a focus on sustainable agricultural practices, will be essential for ensuring food security and preserving plant biodiversity for future generations. The integration of traditional knowledge with modern genomics holds the key to unlocking the full potential of plant genetic resources.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Centre of Origin
The geographic region where a plant species first evolved and exhibits the greatest genetic diversity.
Homologous Series
A group of chemical compounds or plant species with similar structures, exhibiting a gradual and predictable change in properties.

Key Statistics

Vavilov identified 12 primary centres of origin, collectively containing approximately 70% of the world's major crop species.

Source: Vavilov’s work (knowledge cutoff)

The genome size of *Triticum aestivum* (bread wheat) is approximately 17 Gb, reflecting its complex evolutionary history.

Source: Knowledge cutoff

Examples

Wild Rice (Oryza rufipogon)

Serves as a crucial gene pool for improving cultivated rice (*Oryza sativa*), providing resistance to diseases like bacterial blight and tolerance to abiotic stresses.

Potato (Solanum tuberosum)

Wild potato relatives found in the Andes Mountains possess genes for resistance to potato late blight, a devastating disease.

Frequently Asked Questions

Why are centres of origin important for crop improvement?

Centres of origin hold a wealth of genetic diversity, including genes for disease resistance, drought tolerance, and improved yield, which can be introduced into cultivated crops.

How does the Law of Homologous Series apply to plant breeding?

It helps breeders understand the predictable patterns of variation within plant families, enabling them to select for desired traits and develop improved varieties.

Topics Covered

BotanyGeneticsEvolutionPlant EvolutionGenetic VariationCrop Origins