UPSC MainsAGRICULTURE-PAPER-II202420 Marks
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Q27.

What is phytochrome? Discuss its two forms and differentiate them.

How to Approach

This question requires a detailed explanation of phytochrome, its two forms, and their differences. The approach should begin by defining phytochrome and its role in plant development. Subsequently, Pr and Pfr forms should be described, focusing on their chemical structures, conversion processes, and physiological effects. A comparative table highlighting the key differences will enhance clarity. Finally, the significance of phytochrome in agriculture and plant science should be briefly mentioned. Structure: Definition -> Pr form -> Pfr form -> Comparison Table -> Significance.

Model Answer

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Introduction

Phytochrome is a photoreceptor protein found in plants, crucial for mediating various developmental processes in response to light signals. Discovered in the 1950s, it plays a pivotal role in photomorphogenesis, flowering, seed germination, and shade avoidance responses. The understanding of phytochrome's function has revolutionized our understanding of plant-environment interactions, impacting agricultural practices and plant breeding strategies. Recent advances in molecular biology have further elucidated the complex signaling pathways downstream of phytochrome, revealing intricate mechanisms of light perception. This answer will delve into the two forms of phytochrome and their distinguishing characteristics.

What is Phytochrome?

Phytochrome is a pigment found in plant cells that absorbs red and far-red light. It exists in two interconvertible forms, Pr (red-absorbing) and Pfr (far-red-absorbing), and its ratio determines plant responses to light quality and duration. The molecule is a chromoprotein, consisting of a flavoprotein (redox-active) and a protein moiety. It's essential for plants to sense and respond to the changing light conditions of their environment, optimizing growth and survival.

Phytochrome in its Pr Form

Pr (Physiological Red) is the inactive form of phytochrome. It primarily absorbs red light at a wavelength of 660 nm. The absorption of red light by Pr triggers a conformational change, converting it into Pfr. Pr has a relatively low energy state and is the predominant form in the dark or under far-red light conditions. It's relatively stable in the absence of light.

  • Absorption Spectrum: Primarily absorbs red light (660 nm)
  • Conformation: Less active conformation
  • Predominance: Dominant form in darkness or under far-red light
  • Stability: Relatively stable

Phytochrome in its Pfr Form

Pfr (Physiological Far-red) is the active form of phytochrome. It absorbs far-red light at a wavelength of 730 nm. After absorption of far-red light, Pfr reverts to Pr. Pfr is the biologically active form, triggering signaling cascades that influence plant development. In sunlight, Pfr is the more abundant form due to the greater proportion of red light. Pfr can also be converted back to Pr through thermal reversion (a slow process) or through the action of enzymes.

  • Absorption Spectrum: Primarily absorbs far-red light (730 nm)
  • Conformation: More active conformation
  • Predominance: Dominant form under red light or in sunlight
  • Stability: Less stable; undergoes thermal reversion to Pr

Comparison of Pr and Pfr

The following table summarizes the key differences between Pr and Pfr:

Feature Pr (Red-absorbing) Pfr (Far-red-absorbing)
Absorption Wavelength (nm) 660 730
Activity Inactive Active
Conversion Trigger Red light Far-red light
Stability More stable Less stable; undergoes thermal reversion
Predominance (Sunlight) Lower Higher

Significance of Phytochrome

Phytochrome plays a crucial role in several aspects of plant life. It influences seed germination, stem elongation, leaf expansion, flowering time, and pigment synthesis. Understanding phytochrome’s function is vital for optimizing crop yields and developing strategies to enhance plant resilience to varying light conditions. The concept of phytochrome is applied in controlled environment agriculture to manipulate light spectra and optimize plant growth. For instance, LED lighting systems are designed to adjust red and far-red ratios to promote desired plant characteristics.

Conclusion

In conclusion, phytochrome is a vital photoreceptor protein that enables plants to perceive and respond to light signals. The interconversion between its Pr and Pfr forms, driven by red and far-red light respectively, governs a wide range of developmental processes. Continued research into phytochrome signaling pathways promises to further enhance our ability to manipulate plant growth and improve agricultural productivity, especially in the context of changing climates and resource constraints. The understanding of this fascinating molecule underscores the intricate interplay between plants and their environment.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Photomorphogenesis
The developmental changes in plants in response to light signals, beyond the effects of photosynthesis.
Thermal Reversion
The conversion of Pfr back to Pr due to heat, a relatively slow process.

Key Statistics

Approximately 90% of the light absorbed by plants is red or far-red light. (Source: Knowledge Cutoff)

Source: General knowledge of plant physiology

The ratio of Pfr to Pr can change rapidly based on light quality, sometimes within minutes. (Source: Knowledge Cutoff)

Source: General knowledge of plant physiology

Examples

Shade Avoidance Response

When a plant is shaded by another, the ratio of Pfr to Pr decreases, triggering stem elongation to reach sunlight. This is a crucial survival mechanism in dense plant communities.

Flowering Time Regulation

The Pfr/Pr ratio influences flowering time in many plant species, often acting as a signal of day length (photoperiod).

Frequently Asked Questions

Why are there two forms of phytochrome?

The two forms allow plants to sense not only the presence of light but also the *quality* of light. The ratio of red to far-red light provides information about the plant's environment, such as whether it is in direct sunlight or shaded by other plants.

Can phytochrome be affected by factors other than light?

Yes, phytochrome can be influenced by factors like temperature, nutrient availability, and interactions with other plant hormones.

Topics Covered

BotanyPlant PhysiologyPlant HormonesPhotomorphogenesisLight Signaling