UPSC MainsBOTANY-PAPER-II201410 Marks
हिंदी में पढ़ें
Q19.

Mechanism of drought resistance

How to Approach

This question requires a detailed understanding of the physiological and morphological adaptations plants employ to survive drought conditions. The answer should be structured around the different mechanisms – avoidance, tolerance, and resilience – with specific examples. Focus on both constitutive (always present) and induced (developed in response to stress) mechanisms. Mentioning the hormonal regulation involved will add depth. A clear, organized presentation with relevant examples is crucial for a good score.

Model Answer

0 min read

Introduction

Drought, a major abiotic stress, significantly impacts plant growth, productivity, and survival globally. It’s projected to become more frequent and severe with climate change, posing a substantial threat to food security. Plants have evolved a remarkable array of mechanisms to cope with water scarcity, broadly categorized into drought escape, drought avoidance, and drought tolerance. These mechanisms involve morphological, physiological, and biochemical adaptations that allow plants to survive, reproduce, and maintain function under water-limited conditions. Understanding these mechanisms is crucial for developing drought-resistant crops.

Drought Resistance Mechanisms

Drought resistance isn’t a single trait but a complex interplay of various adaptive strategies. These can be broadly classified into three main categories:

1. Drought Avoidance

Drought avoidance strategies allow plants to maintain a high water potential despite reduced water availability. This is achieved by:

  • Reduced Water Loss:
    • Stomatal Regulation: Plants can close their stomata to reduce transpiration. This is often mediated by abscisic acid (ABA), a plant hormone synthesized under drought stress.
    • Leaf Modifications: Smaller leaf size, thicker cuticle, presence of trichomes (leaf hairs), and sunken stomata reduce water loss. Example: Oleander (Nerium oleander) has sunken stomata and a thick cuticle.
    • Leaf Rolling/Folding: Reduces the surface area exposed to sunlight and wind, minimizing transpiration. Example: Many grasses exhibit leaf rolling.
  • Enhanced Water Uptake:
    • Deep Root Systems: Allow access to water deeper in the soil profile. Example: Mesquite trees (Prosopis spp.) have extremely deep root systems.
    • Increased Root Surface Area: Root hairs and lateral roots increase the absorptive capacity.

2. Drought Tolerance

Drought tolerance mechanisms allow plants to survive and function even when fully dehydrated. These include:

  • Osmotic Adjustment: Accumulation of compatible solutes (proline, glycine betaine, sugars) in the cytoplasm lowers the osmotic potential, allowing cells to maintain turgor and continue metabolic processes.
  • Desiccation Tolerance: The ability to withstand extreme dehydration without irreversible damage. Example: Resurrection plants (Selaginella lepidophylla) can survive almost complete desiccation for years and revive upon rehydration.
  • Antioxidant Defense System: Drought stress induces the production of reactive oxygen species (ROS). Plants enhance their antioxidant enzyme systems (superoxide dismutase, catalase, peroxidase) to scavenge ROS and protect cellular components.
  • Protective Proteins: Synthesis of LEA (Late Embryogenesis Abundant) proteins, which protect cellular structures during dehydration.

3. Drought Resilience

Drought resilience refers to the ability of plants to recover quickly after a drought event. This involves:

  • Rapid Rehydration: Efficient water uptake and re-establishment of turgor pressure.
  • Metabolic Recovery: Restoration of photosynthetic capacity and other metabolic processes.
  • Regrowth and Reproduction: Ability to resume growth and reproduction after drought stress.

Hormonal Regulation

Several plant hormones play crucial roles in drought response:

  • Abscisic Acid (ABA): The primary hormone involved in drought stress signaling. It induces stomatal closure, promotes root growth, and regulates the expression of drought-responsive genes.
  • Auxins: Influence root development and water uptake.
  • Cytokinins: Regulate leaf senescence and delay drought-induced damage.
  • Ethylene: Can promote or inhibit drought tolerance depending on the plant species and stress severity.

Genetic Engineering Approaches

Modern biotechnology offers opportunities to enhance drought resistance in crops through genetic engineering. This includes:

  • Overexpression of ABA biosynthesis genes.
  • Transfer of genes encoding osmoprotectants.
  • Manipulation of stomatal density and regulation.

Conclusion

Drought resistance is a multifaceted phenomenon involving a complex interplay of morphological, physiological, and biochemical adaptations. Understanding these mechanisms is vital for developing strategies to mitigate the impacts of drought on agriculture and ecosystems. Future research should focus on identifying and utilizing genes that confer drought resilience, alongside sustainable water management practices, to ensure food security in a changing climate. Integrating traditional breeding techniques with modern biotechnological approaches holds the greatest promise for developing drought-resistant crops.

Answer Length

This is a comprehensive model answer for learning purposes and may exceed the word limit. In the exam, always adhere to the prescribed word count.

Additional Resources

Key Definitions

Osmoprotectants
Compatible solutes accumulated by plants under drought stress to maintain turgor pressure and protect cellular structures. Examples include proline, glycine betaine, and sugars.
Reactive Oxygen Species (ROS)
Highly reactive molecules generated during drought stress that can cause oxidative damage to cellular components. Plants employ antioxidant defense systems to neutralize ROS.

Key Statistics

Approximately 40% of global land area is classified as arid or semi-arid, impacting the livelihoods of over 2 billion people (UNCCD, 2018 - knowledge cutoff).

Source: United Nations Convention to Combat Desertification (UNCCD)

Global crop production could decline by up to 30% by 2050 due to climate change, with drought being a major contributing factor (IPCC, 2021 - knowledge cutoff).

Source: Intergovernmental Panel on Climate Change (IPCC)

Examples

Cactus Adaptations

Cacti exhibit numerous drought adaptations, including succulent stems for water storage, reduced leaves (spines) to minimize transpiration, and a Crassulacean Acid Metabolism (CAM) photosynthetic pathway, which allows them to open stomata at night to reduce water loss.

Frequently Asked Questions

What is the difference between drought resistance and drought tolerance?

Drought resistance refers to the plant's ability to avoid drought stress altogether, while drought tolerance refers to its ability to survive and function despite experiencing drought stress. Resistance is proactive, while tolerance is reactive.

Topics Covered

BiologyPlant SciencePlant PhysiologyStress ResponseAdaptation